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Allen F. Sanborn Canada U.S.A.

Two new cicada publications worth reading

Two relatively new cicada publications that should be worth reading:

1) Avian Predation Pressure as a Potential Driver of Periodical Cicada Cycle Length by Walter D. Koenig and Andrew M. Liebhold, The American Naturalist. This is a newly electronically published paper about what drives the long, prime-numbered lifecycle of Magicicada periodical cicadas.

Abstract:

The extraordinarily long life cycles, synchronous emergences at 13- or 17-year intervals, and complex geographic distribution of periodical cicadas (Magicicada spp.) in eastern North America are a long-standing evolutionary enigma. Although a variety of factors, including satiation of aboveground predators and avoidance of interbrood hybridization, have been hypothesized to shape the evolution of this system, no empirical support for these mechanisms has previously been reported, beyond the observation that bird predation can extirpate small, experimentally mistimed emergences. Here we show that periodical cicada emergences appear to set populations of potential avian predators on numerical trajectories that result in significantly lower potential predation pressure during the subsequent emergence. This result provides new support for the importance of predators in shaping periodical cicada life history, offers an ecological rationale for why emergences are synchronized at the observed multiyear intervals, and may explain some of the developmental plasticity observed in these unique insects.

Order it from JSTOR.

2) The Cicadas (Hemiptera: Cicadoidea: Cicadidae) of N. America North of Mexico by Allen F. Sanborn and Maxine S. Heath. 227 pages.

A comprehensive review of the North Amerian cicada fauna that provides information on synonymies, type localities, and type material. There are 170 species and 21 subspecies found in continental N. America north of Mexico. The book has 211 figures with each species photographed in color.

Buy it from the Entomological Society of America website. I’ve already ordered mine.

I can’t wait for Sanborn’s book on Central and South America (hopefully, that will arrive within the next few years).

Update:

I’ve received Allen F. Sanborn and Maxine S. Heath’s book. It’s focus is on identifying all species of cicada fauna in North America, north of Mexico, as the title says. It also identifies species that were reported to exist in this location, but do not. The book provides maps and common attributes of each genus of cicada, and then for each species it provides photos of the holotype (and the location of the holotype), as well as a history of its taxonomy.

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Chremistica Dundubia Huechys Purana Singapore

Learn about the cicadas of Singapore

Want to learn more about the cicadas of Singapore?

The National University of Singapore has six PDF documents about six species of cicadas living in Singapore. Each document contains photos of cicadas, and plenty of interesting information.

1) Record Of The Cicada, Purana usnani Duffels & Schouten In Singapore, With Preliminary Acoustic Analysis by Tzi Ming Leong (Nature In Singapore 2012 5: 13—17 Date Of Publication: 17 January 2012). Link: rmbr.nus.edu.sg/nis/bulletin2012/2012nis013-017.pdf.

This document features photos of the P. usnani as well as analysis of their songs.

A video of Purana usnani singing:

2) Oviposition By The Black And Scarlet Cicada, Huechys sanguinea (De Geer, 1773) In Singapore by Tzi Ming Leong and Ali bin Ibrahim (Nature In Singapore 2011 4: 303—306 Date Of Publication: 18 October 2011). Link: rmbr.nus.edu.sg/nis/bulletin2011/2011nis303-306.pdf.

Huechys sanguinea is a small but beautiful black and red cicada. This document features photos of an H. sanguinea ovipositing (laying eggs) in a tree branch.

Huechys sanguinea
An image of H. sanguinea, which can be found in Singapore, Thailand and other S.E. Asian countries (and often in acrylic keychains on ebay).

3) Records Of The Cicada, Chremistica umbrosa (Distant, 1904) In Singapore, With Accounts Of Its Mass Emergence by Tzi Ming Leong, Aminurashid and Benjamin P. Y-H. Lee (Nature In Singapore 2011 4: 163—175 Date Of Publication: 15 June 2011). Link: rmbr.nus.edu.sg/nis/bulletin2011/2011nis163-175.pdf.

This document features information about distribution, emergence, bio-acoustics, communal feeding, and predation.

Here’s a video of an aggregation C. umbrosa urinating and singing:

4) Records Of The Black And Golden Cicada, Huechys fusca Distant, 1892 In Singapore, With Natural History Observations by Tzi Ming Leong, Mishak Shunari, Laurence Y. K. Leong, and Sai Khoon Foo (Nature In Singapore 2011 4: 203—211 Date Of Publication: 8 July 2011). Link: rmbr.nus.edu.sg/nis/bulletin2011/2011nis203-211.pdf.

This document features information about emergence, bio-acoustics, mating, and oviposition of Huechys fusca.

A video of Huechys fusca singing:

5) Records Of The Black And Scarlet Cicada, Huechys sanguinea (De Geer) In Singapore, With Notes On Its Emergence by Ali bin Ibrahim and T. M. Leong (Nature In Singapore 2009 2: 317—322 Date Of Publication: 5 August 2009). Link: rmbr.nus.edu.sg/nis/bulletin2009/2009nis317-322.pdf.

This document features observations of Huechys sanguinea. Huechys sanguinea is a beautiful cicada.

6) The Jade-Green Cicada, Dundubia vaginata (Fabricius, 1787) In Singapore, With Notes On Emergence, Bioacoustics, And Mating by Tzi Ming Leong, Mishak Shunari, Aminurashid and Timothy D. Harvey-Samuel (Nature In Singapore 2011 4: 193—202 Date Of Publication: 5 July 2011). Link: rmbr.nus.edu.sg/nis/bulletin2011/2011nis193-202.pdf.

This document features information about emergence, bio-acoustics, and mating of Dundubia vaginata. Dundubia are known for there huge opercula (the structures on their abdomen that cover the cicada’s tympanum (tympanum are a cicada’s ear drums).

Categories
Anatomy Australia

Blue Cicadas

Blue cicadas. Did you know they exist? They do… at least in Australia.

What’s That Bug recently posted a photo of a blue Bladder Cicada from Australia (Cystosoma saundersii). It’s a great find. Cystosoma saundersii are typically green.

Then there is the Blue Moon blue colored morph of Cyclochila australasiae:

Cyclochila australasiae, Blue Moon, by David Emery
Photo by David Emery

Cyclochila australasiae come in many colors, but the most common color is green. “Blue Moon” is a good nickname for these cicadas because they are rare and only found, idiomatically speaking, “once in a Blue Moon”.

So, why are some cicadas blue, when their species is typically green? Here is a quote from the paper Blue, red, and yellow insects by B. G. BENNETT, Entomology Division, DSIR, Private Bag, Auckland, New Zealand:

The colours of insects are often due to a complex mixture of pigments, some of which
are concentrated from their diet. These are carotenoids, flavonoids, and anthraquinones, and some are porphyrins made from the breakdown of plant chlorophyll. Insectoverdin is a common green pigment produced by a mixture of blue and yellow compounds. The blue is tetrapyrrole, but sometimes an anthocyanin, and the yellow is a carotenoid.

Blue + yellow = green. If the yellow is missing, you get a blue cicada. I heard that, at least in the case of the Cyclochila australasiae, the blue cicadas are typically females. Perhaps something related to genetics or behavior of the females leads to an inability to process the caroteniods ingested along with their diet (tree fluids). I’m not sure, but it’s a topic that fascinates me, so I’ll continue to look into it.

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Cystosoma David Emery Mating

Mating Bladder cicadas



Mating bladders 2, originally uploaded by ozzicada.

An excellent photo of mating Bladder cicadas (Cystosoma saundersii) by David Emery.

Categories
Cacama Tacuini (Cryptotympanini)

Cacama aka Cactus Dodgers

Cacama valvata

A Cacama moorei (female) photo taken by Adam Fleishman.

Cacama is a genus of cicadas, known as Cactus Dodgers, found in Mexico and Southwestern United States. They are known for their affinity for cacti like prickly pear & cholla, and are most likely named Cactus Dodgers for their ability dodge the needles of their favorite plants. They are primarily black, gray, white, and beige colored; well camouflaged for the desert.

The two most common species seem to be Cacama moorei and Cacama valvata. The best way to tell them apart is C. moorei have a lot of orange on their ventral side.

Eyes Gray to golden beige
Ventral side Mostly white, heavy pruinose
Legs Black to golden beige
Dorsal side Gray to Black, with rust, golden or beige highlights. Prominent white pruinose along the sides of the mesonotum, and the 1st tergite (dorsal abdominal segment) of the abdomen.
Wings hyaline, with black to golden beige viens

According to BugGuide there are 12 species of Cacama: C. californica , C. carbonaria, C. crepitans, C. collinaplaga, C. dissimilis, C. furcata, C. longirostris, C. maura, C. moorei, C. pygmaea, C. valvata and C. variegata.

Cacama was the lord of the Aztec kingdom of Tezcuco (see The History of the Conquest of Mexico, by W.H. Prescott), who met his end at the hands of Spanish conquistadors. Cacama lives on in these winged desert treasures.

Some Cacama links:

The Insect Singers website has the song of a Cacama valvata.

A nice photo of a Cacama furcata taken in New Mexico.


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Australia

Great website: The cicadas of central eastern Australia

If you are located in Australia and like cicadas, you should visit The cicadas of central eastern Australia, a website created by Lindsay Popple.

Popple’s website includes: photos, maps, range & season, habits, and recordings of the song of dozens of Australian cicadas. Very complete and well done.

The cicadas of central eastern Australia

The site was recommended to me by David Emery.

Categories
Chimneys

Cicada Turrets or Chimneys

Cicada nymphs create what we call turrets or chimneys above the tunnel hole where they will eventually emerge. These chimneys are made from soil. They are a positive indication of where cicada nymphs are underground and that they will emerge soon. Cicadas seem to build chimneys in wetter, muddier areas; in dry areas they will simply make holes at the surface. The chimneys will help keep water and mud from back-filling their holes, so they can continue to breathe, take a peek out and prepare to emerge.

Magicicada Tunnel (by Les Daniels):

Magicicada chimney
Magicicada chimney

A cicada tunnel in Bangkok Thailand (by Santisuk Vibul):
cicada tunnel view from top Thailand photo by Santisuk Vibul

Three Magicicada chimneys (by Roy Troutman):

Magicicada nymph turret by Roy:

This is video taken of a magicicada nymph turret. These turrets are often found in wet areas.

Magicicada nymph turret from Roy Troutman on Vimeo.

Categories
Brood II Magicicada Periodical

Getting Ready for the 2013 Brood II Emergence

Brood II will next emerge in 2030.

This page has not been updated since 2013.

Cicada Mania was started back in 1996, the last time Brood II emerged! The Spring of 2013 will be our first chance to see the children of the cicadas that emerged 17 years ago. Here is the basic information you need to know about the 2013 Brood II emergence.

Even though the emergence is 5 to 6 months away, it is never too early to begin planning… especially if you are a cicada maniac like me.

Magicicada septendecimThere will be plenty of cicadas on hand.

When will the Brood II cicadas emerge?

Brood II cicadas will emerge sometime in the Spring of 2013. They typically emerge once the soil 8 inches (20cm) below the surface gets to 64 degrees Fahrenheit (18º C). If we have a hot Spring, as we did in 2012, the cicadas could emerge in mid-to-late April. If we have a moderate Spring, the cicadas will wait until May.

Where will they emerge?

Brood II will emerge in parts of Connecticut, Maryland, North Carolina, New Jersey, New York, Pennsylvania, and Virginia.

Magicicadas won’t emerge everywhere in the states mentioned above. They might not exist in your town or neighborhood (particularly if trees were removed from your neighborhood).

More information about where the cicadas will emerge.

What is a Magicicada cicada?

Magicicada is a genus of periodical cicadas known for emerging in massive numbers in 17 or 13 year cycles/periods. The cicadas emerging in 2013 have 17 year life-cycles. Magicicada are also organized into broods. There are 12 broods of 17 year cicadas, and the brood emerging in 2013 is Brood II (Brood Two).

There are 3 species of 17-year Magicicada: M. septendecim (aka “decims”), M. cassini, and M. septendecula. The adults of all three species have black bodies with orange markings, and almost all have red-orange eyes (some have white or multi-colored eyes.

Here is some video and audio of 17-year Magicicada. This will give you an idea of what to expect:

Cicada Mania, best of 2007, part 1 by Dan from Cicada Mania on Vimeo.

More information:


1907 Map Marlatt, C.L.. 1907. The periodical cicada. Washington, D.C. : U.S. Dept. of Agriculture, Bureau of Entomology.

Marlatt 1907 02 Brood II

Categories
Ariasa Brazil Fidicina Fidicinini Fidicinoides

Brazil Cicada Identification Challenge, Part 2

Here is part 2. Jairo of Cigarras do Brasil — Brazilian Cicadas asked for our help to identify some unknown cicada species from Brazil. The following photographs feature cicadas from Brazil we want to identify. We are hoping folks in the cicada research community can help.

Note: All of these cicadas were photographed at Paraibuna, São Paulo. This town is close to the Paraíba Valley (Vale do Paraíba), and to São José dos Campos and Caçapava.

1) Possibly an Ariasa sp. (about 1/2 inch (1,3cm))

Possibly an Ariasa sp. (about 1/2 inch (1,3cm))

2) Possibly a Quesada sodalis (about 1 1/2 inch (4cm))

Possibly a Quesada sodalis (about 1 1/2 inch (4cm))

Possibly a Quesada sodalis (about 1 1/2 inch (4cm))

Cicadas 3, 4 and 5 were identified by David Emery as Fidicinoides picea (Walker, 1850). Learn more about Fidicnoides picea.

3) Identified: Fidicinoides picea.

Fidicinoides picea

4) Identified: Fidicinoides picea.

Fidicinoides picea

5) Identified: Fidicinoides picea.

Fidicinoides picea

Categories
Allen F. Sanborn FAQs Zammara

How to learn more about cicadas, by searching for cicada researchers

Just about anything and everything ends up on the web these days, including research papers written by cicada researchers. Many of these papers are easily downloaded from the web, and once downloaded you can read them and expand your knowledge about cicadas.

This weekend I was looking for information about cicadas from Central and South America (the Neotropic ecozone). Allen F. Sanborn, Ph.D is well known for his research of cicadas of that region, so I searched for some of his research papers. Google will retrieve all PDF (Adobe Acrobat) files that contain the word cicada and the name Allen F. Sanborn, when you search for “Allen F. Sanborn cicada ext:pdf” (remove the quotes when you search).

Some interesting papers I found include:

Checklist of the cicadas (Insecta: Hemiptera: Cicadidae) of Paraguay including new records for six species (Sanborn, Allen F., 2011). This paper includes a long list of cicada species, which I used to look for images of cicadas on Flickr.com.

The new records increase the known cicada diversity 37.5% bringing the total number of cicada species reported in Paraguay to 22 species in 12 genera representing five tribes and three subfamilies of the family Cicadidae. There are currently no known endemic species.

Two New Zammara Species from South America (Hemiptera: Cicadomorpha: Cicadidae) (Sanborn, Allen F., Florida Entomologist 87(3),2004). This paper includes many photographs, which make cicada identification easy.

ABSTRACT
Two new members of the widespread Neotropical genus Zammara Amyot & Serville, Zammara
olivacea n.sp. from Columbia and Zammara medialinea n.sp. from Venezuela are described.

Key Words: new species, taxonomy, cicada, Zammara, Columbia, Venezuela.

New Records for the Cicada Fauna from Four Central American Countries (HEMIPTERA: CICADOIDEA: CICADIDAE (Sanborn, Allen F.; Florida Entomologist 89(1), 2006). This article features a map with cicada species names.

ABSTRACT
Analysis of museum specimens has added to the cicada fauna of Belize, El Salvador, Guate- mala, and Honduras. Information on the cicada fauna reported in the literature as well as the first records of cicada species to the fauna are reported here to provide a more accurate un- derstanding of cicada diversity in each country and the region. The new records represent an increase of 75, 14, 110, and 320%, respectively, to the cicada faunal diversity of each country.
Key Words: cicadas, biodiversity, Central America

If you use my Google formula, you can find these papers too.

The The Current Status of Cicada Taxonomy on a Region-by-Region Basis page on Cicada Central is a good resource for learning about other cicada researchers.