Cicada Mania

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April 18, 2020

Davis’ Key to Species of the Genus Tibicen found in the Southeastern United States

I’ve been updating the North American cicada pages on this site, and I’ve been including snippets (and some chunks) from William T. Davis’ cicada papers from the early 1900s. They’re in the public domain and excellent resources.

Davis provided a key of cicadas that belong to the then genus Tibicen in his 1918 article Mississippi Cicadas, with a Key to the Species of the Southeastern United States from volume 26 of the Journal of The New York Entomological Society. Download it from archive.org.

Mississippi Cicadas

Since 1918, genus and some species names have changed, so I’m going to present the key here, with highlighted notes on the updated names + images (when I have them). I’ll try to replicate the formatting of the original document as best I can.

Here goes…

Key to Species of the Genus Tibicen found in the Southeastern United States.

Note: the cicadas in the key are now organized in three genera: Neotibicen (A B), Megatibicen (A BB), and Diceroprocta (AA).

A. Large, heavy bodied species; head broad, uncus simple, and first cross vein in the fore wings starting from radius 3 far back, or about one third distant from base of first marginal cell.

     B. Uncus longer than broad. Black species with green or greenish markings and black area on the central part of the abdomen beneath, except in sayi [sayi = Neotibicen tibicen tibicen], and new variety of davisi [new variety of davisi = Neotibicen davisi harnedi].

Note: this group of cicadas (B) are now organized under the genus Neotibicen, not Tibicen.

         C. Hind margin of pronotum or collar, green or greenish.

             A narrow irregular area of black on the under side of the abdomen; opercula short and broad, and usually in the males an attenuated, pruinose stripe each side on the dorsum of segment three … pruinosa (Say). [pruinosa = Neotibicen pruinosus pruinosus].

N. pruinosus

Neotibicen pruinosus pruinosus
photo by Paul Krombholz.

             Dorsum of abdomen with the hind margin of the segments more or less brown and generally but a trace of pruinose stripe each side on segment three … pruinosa var. winnemanna (Davis) [pruinosa var. winnemanna = Neotibicen winnemanna].

Neotibicen winnemanna Garner NC
Photo of a Neotibicen winnemanna.

             Dorsum of abdomen shining black with a broad pruinose mark each side on segment three; blackened area on under side of abdomen more in the nature of an even stripe … pruinosa var latifasciata (Davis) [pruinosa var. latifasciata = Neotibicen latifasciatus].

             A longitudinal band of black on the under side of the abdomen, the opercula more lobate, and the margin of the front wings suddenly bent near the middle … linnei (Smith & Grossbeck) [linnei = Neotibicen linnei].

Neotibicen linnei
Photo of a Neotibicen linnei.

             A definite longitudinal band of black on the under side of the abdomen; head with the front rather prominent. Not a large species … canicularis (Harris) [canicularis = Neotibicen canicularis].

             An irregular band of black on the under side of the abdomen, head rounded in front; a rather small species … davisi (Smith & Grossbeck) [davisi = Neotibicen davisi davisi].

             Abdomen greenish centrally on under side, blackened area wanting, marginal cells of fore wings clouded … davisi var. harnedi new variety [davisi var. harnedi = Neotibicen davisi harnedi].

Krombholz Davisi compared
Photo by Paul Krombhold. Neotibicen davisi harnedi (left), Neotibicen davisi davisi (right).

        CC. Hind margin of pronotum or collar black or nearly so (except in sayi var. australis).

            D. Central area of the abdomen beneath black.

                 Opercula long and with the legs usually somewhat chest- nut colored ; the uncus when seen in profile forked, resembling the open mouth of a snake … similaris (Smith & Grossbeck) [similaris = Neotibicen similaris similaris].

                 Opercula much shorter, more rounded, and the black area on the under side of the abdomen in the nature of an even stripe. Uncus not forked … lyricen (De Geer) [lyricen = Neotibicen lyricen lyricen].

                 Blacker than typical lyricen, lacking the considerable amount of fulvous markings on the pronotum and mesonotum. A fulvous somewhat anchor-shaped mark centrally on the pronotum … lyricen var. engelhardti (Davis) [lyricen var. engelhardti = Neotibicen lyricen engelhardti].

            DD. Central area of the abdomen not black beneath, often pruinose, as well as the long opercula.

                 Collar black, often with a greenish spot each side near the outer angles. … sayi (Smith & Grossbeck) [sayi = Neotibicen tibicen tibicen].

                 Collar all green or nearly so, as well as the pronotum and mesonotum … sayi var. australis (Davis) [sayi = Neotibicen tibicen australis].

    BB. Uncus broad at the base, triangular in shape and generally about as broad as long. Opercula broad and rounded at the extremities no definite black area on the central part of the abdomen beneath, usually unicolorus.

Note: this group of cicadas (BB) are now organized under the genus Megatibicen, not Tibicen.

         E. Wings long and narrow, collar 2 mm. or less in breadth at central portions ; dorsum of abdomen black or nearly so.

             Basal cell of fore wings rusty in color, anal cells (mem- branes) of both pair of wings gray; usually expands no mm. or more … resonans (Walker) [resonans = Megatibicen resonans].

Neotibicen resonans photos by Joe Green from 2007, taken in Florida.
Photo of a Megatibicen resonans by Joe Green.

             Basal cell of fore wings often black or nearly so, anal cells of both pair of wings yellowish. Expands about 100 mm … figurata (Walker) [figurata = Megatibicen figuratus].

M. figurata
Photo of a Megatibicen figuratus by Paul Krombholz.

         EE. Wings broad, hind margin of the pronotum or collar green or greenish and more than 2 mm. broad.

             F. Anal cells or membranes at base of fore and hind wings gray.

                 Dorsal segments of the abdomen not margined with brown ; in fresh specimens the basal segments pruinose, also the terminal segments, leaving the four middle segments black. A large species expanding over no
mm. … auletes (Germar) [auletes = Megatibicen auletes].

Megatibicen auletes, the largest cicada in North America
Photo of a Megatibicen auletes.

             FF. Anal cells or membranes at base of fore and hind wings light orange, two prominent marks on the mesonotum resembling the Hebrew letter resh inverted.

                 Fore wings with the first and second cross veins clouded, and the dorsum of the abdomen brownish or brownish black … resh (Haldeman) [resh = Megatibicen resh].

Megatibicen resh molting adult
Photo of a Megatibicen resh.

                 Fore wings with the first and second cross veins but faintly or not at all clouded and the abdominal segments margined posteriorly with brown. In fresh specimens there is usually a median row of white spots on the dorsum of the abdomen … marginalis (Walker) [marginalis = Megatibicen pronotalis walkeri].

Megatibicen pronotalis photo by Roy Troutman, taken in Batavia, Ohio
Photo of a Megatibicen pronotalis walkeri by Roy Troutman.

AA. Small species ; wings starting from about the middle of the first marginal cell.

Note: this group of cicadas (AA) are now organized under the genus Diceroprocta, not Tibicen.

     G. First and second cross veins of fore wings clouded.

         Expanse of wings about 90 mm … biconica (Walker) [biconica = Diceroprocta biconica].

         Expanse of wings about 60 mm … olympusa (Walker) [olympusa = Diceroprocta olympusa].

Diceroprocta olympusa photos by Joe Green from 2007.
Photo of a Diceroprocta olympusa by Joe Green.

     GG. First and second cross veins of fore wings not clouded, wings clear throughout and expanding about 70 mm.

         Head rather large, front rounded, collar greenish or yellowish and contrasted in color rather sharply with the brown and black of pronotum and mesonotum … viridifascia (Walker) [viridifascia = Diceroprocta viridifascia].

         Head proportionately smaller than in the last ; front more pro- truding;- collar not so contrastingly colored and fore wings narrower … vitripennis (Say) [vitripennis = Diceroprocta vitripennis].


Diceroprocta vitripennis photo by Paul Krombholz.

and that’s all folks…

April 13, 2020

Megatibicen pronotalis pronotalis Davis, 1938

Megatibicen pronotalis pronotalis Davis, 1938

Image

Megatibicen pronotalis pronotalis Davis, 1938
Photo credit: by Dan Mozgai (Bill Reynolds collection)

Song type: Call


Source: ©Insect Singers | Species: M. pronotalis pronotalis

Video Playlist

Playlists contain multiple videos found on YouTube.

Name, Location and Description

Classification:

Family: Cicadidae
Subfamily: Cicadinae
Tribe: Cryptotympanini
Subtribe: Cryptotympanina
Genus: Megatibicen
Species: Megatibicen pronotalis pronotalis Davis, 1938
Species: Megatibicen pronotalis walkeri Metcalf, 1955 aka Walker’s Cicada

List of sources

  1. Full Binomial Names: ITIS.gov
  2. Common names: BugGuide.net; The Songs of Insects by Lang Elliott and Wil Herschberger; personal memory.
  3. Locations: Biogeography of the Cicadas (Hemiptera: Cicadidae) of North America, North of Mexico by Allen F. Sanborn and Polly K. Phillips.
  4. Descriptions, Colors: personal observations from specimens or photos from many sources. Descriptions are not perfect, but may be helpful.

Notes:

  • Some descriptions are based on aged specimens which have lost some or a lot of their color.

April 4, 2020

Megatibicen resonans (Walker, 1850) aka Southern Resonant Cicada

Filed under: Cryptotympanini,Megatibicen,United States — Tags: — Dan @ 4:51 pm

Megatibicen resonans (Walker, 1850) aka Great Pine Barrens Cicada.

Neotibicen resonans photos by Joe Green from 2007, taken in Florida.
Photo by Joe Green.

â­˘ All Megatibicen resonans images & information on cicadamania.com.

Song type: Call


Source: ©Insect Singers | Species: M. resonans

Song type: Call


Source: ©Joe Green | Species: M. resonans

Video

Video Playlist

Playlists contain multiple videos found on YouTube.

Name, Location and Description

Key to Species by Wm. T. Davis from MISSISSIPPI CICADAS, WITH A KEY TO THE SPECIES OF THE SOUTHEASTERN UNITED STATES.1

A. Large, heavy bodied species ; head broad, uncus simple, and first cross vein in the fore wings starting from radius 3 far back, or about one third distant from base of first marginal cell.

BB. Uncus broad at the base, triangular in shape and generally about as broad as long. Opercula broad and rounded at the extremities no definite black area on the central part of the abdomen beneath, usually unicolorus.

E. Wings long and narrow, collar 2 mm. or less in breadth at central portions ; dorsum of abdomen black or nearly so.

Basal cell of fore wings rusty in color, anal cells (membranes) of both pair of wings gray; usually expands 110 mm. or more resonans (Walker).

Classification:

Family: Cicadidae
Subfamily: Cicadinae
Tribe: Cryptotympanini
Subtribe: Cryptotympanina
Genus: Megatibicen
Species: Megatibicen resonans (Walker, 1850)

List of sources

  1. Davis, W.T. 1918. MISSISSIPPI CICADAS, WITH A KEY TO THE SPECIES OF THE SOUTHEASTERN UNITED STATES. Journal of the New York Entomological Society, Vol. XXVI, Nos. 3-4. Read it on archive.org.
  2. Full Binomial Names: ITIS.gov
  3. Common names: BugGuide.net; The Songs of Insects by Lang Elliott and Wil Herschberger; personal memory.
  4. Locations: Biogeography of the Cicadas (Hemiptera: Cicadidae) of North America, North of Mexico by Allen F. Sanborn and Polly K. Phillips.
  5. Descriptions, Colors: personal observations from specimens or photos from many sources. Descriptions are not perfect, but may be helpful.

Notes:

  • Some descriptions are based on aged specimens which have lost some or a lot of their color.

Megatibicen resh (Haldeman, 1852) aka Resh Cicada

Filed under: Cryptotympanini,Megatibicen,United States — Tags: — Dan @ 4:30 pm

Megatibicen resh (Haldeman, 1852) aka Resh Cicada.

Megatibicen resh molting adult

â­˘ All Megatibicen resh images & information on cicadamania.com.

Song type: Call


Source: ©Insect Singers | Species: M. resh

Video Playlist

Playlists contain multiple videos found on YouTube.

Name, Location and Description

Key to Species by Wm. T. Davis from MISSISSIPPI CICADAS, WITH A KEY TO THE SPECIES OF THE SOUTHEASTERN UNITED STATES.1

A. Large, heavy bodied species ; head broad, uncus simple, and first cross vein in the fore wings starting from radius 3 far back, or about one third distant from base of first marginal cell.

BB. Uncus broad at the base, triangular in shape and generally about as broad as long. Opercula broad and rounded at the extremities no definite black area on the central part of the abdomen beneath, usually unicolorus.

EE. Wings broad, hind margin of the pronotum or collar green or greenish and more than 2 mm. broad.

FF. Anal cells or membranes at base of fore and hind wings light orange, two prominent marks on the mesonotum resembling the Hebrew letter resh inverted.

Fore wings with the first and second cross veins clouded, and the dorsum of the abdomen brownish or brownish black resh (Haldeman).

Classification:

Family: Cicadidae
Subfamily: Cicadinae
Tribe: Cryptotympanini
Subtribe: Cryptotympanina
Genus: Megatibicen
Species: Megatibicen resh (Haldeman, 1852)

List of sources

  1. Davis, W.T. 1918. MISSISSIPPI CICADAS, WITH A KEY TO THE SPECIES OF THE SOUTHEASTERN UNITED STATES. Journal of the New York Entomological Society, Vol. XXVI, Nos. 3-4. Read it on archive.org.
  2. Full Binomial Names: ITIS.gov
  3. Common names: BugGuide.net; The Songs of Insects by Lang Elliott and Wil Herschberger; personal memory.
  4. Locations: Biogeography of the Cicadas (Hemiptera: Cicadidae) of North America, North of Mexico by Allen F. Sanborn and Polly K. Phillips.
  5. Descriptions, Colors: personal observations from specimens or photos from many sources. Descriptions are not perfect, but may be helpful.

Notes:

  • Some descriptions are based on aged specimens which have lost some or a lot of their color.

Megatibicen pronotalis walkeri Metcalf, 1955 aka Walker’s Cicada

Filed under: Cryptotympanini,Megatibicen,United States — Tags: — Dan @ 3:55 pm

Megatibicen pronotalis walkeri Metcalf, 1955 aka Walker’s Cicada.

Megatibicen pronotalis photo by Roy Troutman, taken in Batavia, Ohio
Photo by Roy Troutman in Batavia, Ohio.

â­˘ All Megatibicen pronotalis images and info on cicadamania.com.

There’s two sub-species of Megatibicen pronotalis: Megatibicen pronotalis pronotalis Davis — found in IA, MO, NE, ND, OK, SD — and Megatibicen pronotalis walkeri Metcalf — found in AL, AR, FL, GA, IL, IN, IA, KS, KY, LA, MD, MI, MN, MS, MO, NE, NC, ND, OH, OK, SD, TN, TX, VA, WV, WI, and WY.

Song type: Call


Source: ©Insect Singers | Species: M. pronotalis walkeri

Video Playlist

Playlists contain multiple videos found on YouTube.

Name, Location and Description

Key to Species by Wm. T. Davis from MISSISSIPPI CICADAS, WITH A KEY TO THE SPECIES OF THE SOUTHEASTERN UNITED STATES.1

A. Large, heavy bodied species ; head broad, uncus simple, and first cross vein in the fore wings starting from radius 3 far back, or about one third distant from base of first marginal cell.

BB. Uncus broad at the base, triangular in shape and generally about as broad as long. Opercula broad and rounded at the extremities no definite black area on the central part of the abdomen beneath, usually unicolorus.

EE. Wings broad, hind margin of the pronotum or collar green or greenish and more than 2 mm. broad.

FF. Anal cells or membranes at base of fore and hind wings light orange, two prominent marks on the mesonotum resembling the Hebrew letter resh inverted.

Fore wings with the first and second cross veins but faintly or not at all clouded and the abdominal segments margined posteriorly with brown. In fresh specimens there is usually a median row of white spots on the dorsum of the abdomen, .marginalis (Walker).

Classification:

Family: Cicadidae
Subfamily: Cicadinae
Tribe: Cryptotympanini
Subtribe: Cryptotympanina
Genus: Megatibicen
Species: Megatibicen pronotalis
Sub-species: Megatibicen pronotalis walkeri Metcalf, 1955
Sub-species: Megatibicen pronotalis pronotalis Davis, 1938

List of sources

  1. Davis, W.T. 1918. MISSISSIPPI CICADAS, WITH A KEY TO THE SPECIES OF THE SOUTHEASTERN UNITED STATES. Journal of the New York Entomological Society, Vol. XXVI, Nos. 3-4. Read it on archive.org.
  2. Full Binomial Names: ITIS.gov
  3. Common names: BugGuide.net; The Songs of Insects by Lang Elliott and Wil Herschberger; personal memory.
  4. Locations: Biogeography of the Cicadas (Hemiptera: Cicadidae) of North America, North of Mexico by Allen F. Sanborn and Polly K. Phillips.
  5. Descriptions, Colors: personal observations from specimens or photos from many sources. Descriptions are not perfect, but may be helpful.

Notes:

  • Some descriptions are based on aged specimens which have lost some or a lot of their color.

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