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June 19, 2018

Z. P. Metcalf Collection of Literature on Auchenorrhyncha

Filed under: Allen F. Sanborn,Papers and Documents — Dan @ 8:58 pm

Here is information about a research resource for Auchenorrhyncha researchers. I’m posting at the request of Lew Dietz and Allen F Sanborn.

Z. P. Metcalf Collection of Literature on Auchenorrhyncha

Dear Colleagues,

We would like to make you aware of an exciting resource that is available to you at the Special Collections Research Center at NCSU Libraries in Raleigh, North Carolina.

The Zeno Payne Metcalf Entomology Research Collection is an outstanding resource for scholars. The link to the finding aid is here:

https://www.lib.ncsu.edu/findingaids/mc00220

And the link to the associated database for individual publications is here: http://metcalf.lib.ncsu.edu/metcalf/

If you would like to make an appointment to view any materials in person, or to request a copy of a publication remotely, please email us at: library_specialcollections@ncsu.edu

We typically need 2-3 business days to pull selected materials from off-site storage. More information about visiting the Special Collections Research Center can be found here: https://www.lib.ncsu.edu/scrc/using-materials

Finally, we are pleased to let you know that low-resolution copies (PDFs) only cost 50 cents per page, but, if you bring a camera (no flash allowed) or cell phone, you can take photographs at no cost in the Special Collections Reading Room. If you are unable to visit in person, we can still provide low-resolution copies (PDFs) to you for the fees outlined above. For security reasons, arrangements to pay using a major credit card (preferred) or bank transfer, should be made by telephone. Copyright law applies; see: https://www.lib.ncsu.edu/scrc/copyright

If you have any further questions about the Metcalf materials or copyright, please don’t hesitate to contact us at library_specialcollections@ncsu.edu

Sincerely, Gwynn Thayer, Acting Department Head, Special Collections Research Center (12 June 2018)

June 2, 2018

New paper on the molecular phylogeny of the cicadas and tribe and subfamily classification

A new paper has been published titled A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily classification by David C. Marshall, Max Moulds, Kathy B. R. Hill, Benjamin W. Price, Elizabeth J. Wade, Christopher L. Owen, Geert Goemans, Kiran Marathe, Vivek Sarkar, John R. Cooley, Allen F. Sanborn, Krushnamegh Kunte, Martin H. Villet, Chris Simon.

The paper was published in Vol 4424, No 1 of Zootaxa. Link to paper.

In a nutshell: These researchers compared the DNA of a variety of cicadas to determine how they are related evolutionarily and how they should be organized in terms of tribes and sub-families.

Abstract:

A molecular phylogeny and a review of family-group classification are presented for 137 species (ca. 125 genera) of the insect family Cicadidae, the true cicadas, plus two species of hairy cicadas (Tettigarctidae) and two outgroup species from Cercopidae. Five genes, two of them mitochondrial, comprise the 4992 base-pair molecular dataset. Maximum-likelihood and Bayesian phylogenetic results are shown, including analyses to address potential base composition bias. Tettigarcta is confirmed as the sister-clade of the Cicadidae and support is found for three subfamilies identified in an earlier morpho- logical cladistic analysis. A set of paraphyletic deep-level clades formed by African genera are together named as Tettigo- myiinae n. stat. Taxonomic reassignments of genera and tribes are made where morphological examination confirms incorrect placements suggested by the molecular tree, and 11 new tribes are defined (Arenopsaltriini n. tribe, Durangonini n. tribe, Katoini n. tribe, Lacetasini n. tribe, Macrotristriini n. tribe, Malagasiini n. tribe, Nelcyndanini n. tribe, Pagi- phorini n. tribe, Pictilini n. tribe, Psaltodini n. tribe, and Selymbriini n. tribe). Tribe Tacuini n. syn. is synonymized with Cryptotympanini, and Tryellina n. syn. is synonymized with an expanded Tribe Lamotialnini. Tribe Hyantiini n. syn. is synonymized with Fidicinini. Tribe Sinosenini is transferred to Cicadinae from Cicadettinae, Cicadatrini is moved to Ci- cadettinae from Cicadinae, and Ydiellini and Tettigomyiini are transferred to Tettigomyiinae n. stat from Cicadettinae. While the subfamily Cicadinae, historically defined by the presence of timbal covers, is weakly supported in the molecular tree, high taxonomic rank is not supported for several earlier clades based on unique morphology associated with sound production.

January 23, 2018

Second edition of Cicadas of North American North of Mexico

Filed under: Allen F. Sanborn,Maxine E. Heath — Dan @ 5:31 am

Allen F. Sanborn and Maxine S. Heath have released the second edition of their book Cicadas of North American North of Mexico.

What’s new:

The second edition includes the addition of four new genera, the removal of two genera, and the addition of a few new species that were described since the first edition. We also added distributional data for all species.

Purchase a copy online from the Entomological Society of America.

Related articles:

November 5, 2017

New paper: thermal adaptation in North American cicadas

Temperature plays an important part in much of cicada behavior, such as determining when they emerge from the ground, and when they are active above ground.

There’s a new paper out from Allen F.Sanborn, James E.Heath, Maxine S.Heath and Polly K.Phillips titled “Thermal adaptation in North American cicadas”.

Here are the highlights:

  • Thermal responses are related specific environments of North America cicadas.
  • Thermoregulatory strategy can influence thermal responses in sympatric species.
  • Emergence time can influence thermal responses in sympatric species.
  • Subspecies in general do not differ in their thermal responses.
  • Thermal responses within a species do not differ in populations separated by more than 7600 km.

And here’s the citation info (even though I’m not citing anything):


Allen F. Sanborn, James E. Heath, Maxine S. Heath, Polly K. Phillips, Thermal adaptation in North American cicadas (Hemiptera: Cicadidae), In Journal of Thermal Biology, Volume 69, 2017, Pages v-xviii, ISSN 0306-4565, https://doi.org/10.1016/j.jtherbio.2017.07.011.
(http://www.sciencedirect.com/science/article/pii/S030645651730181X)

September 17, 2016

What is Megatibicen?!

Update (9/20): I guessed the species correctly: all the Large Flute Players.

Update (9/24): I neglected to note that there’s another paper out there by Young June Lee called Description of three new genera, Paratibicen, Megatibicen, and Ameritibicen, of Cryptotympanini (Hemiptera: Cicadidae) and a key to their species. Link to it here. This manuscript goes beyond one new genera, and instead introduces three: Paratibicen, Megatibicen, and Ameritibicen. Lee’s paper differs from Sanborn & Heath in that the large Neotibicen are spit into Megatibicen and Ameritibicen in Young’s document, but they’re all Megatibicen in Sanborn & Heath’s paper.

Megatibicen

Last night I had a rough night’s sleep. I tossed and turned all night long. I remember looking at the clock and seeing 4am, and thinking “tomorrow is ruined”. Sometime during the night I dreamt of finding thousands of molted Neotibicen exuvia clinging to shrubbery — a rare if not impossible sight in real life.

When I woke I checked my email and found a communication from David Marshall. David is well known and respected in the cicada world for many things including describing the 7th species of Magicicada with John Cooley (link to document), as well as being part of the team who defined the Neotibicen and Hadoa genera (link to paper)1.

David wrote to let me know that Allen F. Sanborn and Maxine S. Heath had published a new paper titled: Megatibicen n. gen., a new North American cicada genus (Hemiptera: Cicadidae: Cicadinae: Cryptotympanini), 2016, Zootaxa Vol 4168, No 3.(link).

So, what is MEGATIBICEN? Assumptions after the abstract.

Here is the abstract:

The genus Tibicen has had a confusing history (see summary in Boulard and Puissant 2014; Marshall and Hill 2014; Sanborn 2014). Boulard and his colleague (Boulard 1984; 1988; 1997; 2001; 2003; Boulard and Puissant 2013; 2014; 2015) have argued for the suppression of Tibicen and the taxa derivatived from it in favor of Lyristes Horváth. Boulard’s argument for suppression was first described in Melville and Sims (1984) who presented the case for suppression to the International Commission on Zoological Nomenclature with further comments made by Hamilton (1985), Boulard (1985), and Lauterer (1985). A lack of action resulted in additional comments being published in 2014 again supporting the retention (Sanborn 2014; Marshall and Hill 2014) or the suppression (Boulard and Puissant 2014) of Tibicen.

My guess, without reading the document, is that Megatibicen includes the larger North America Neotibicen species, including the “auletes group” (N. auletes, N. resh, N. figuratus, N. resonans), the “pronotalis group” (N. pronotalis, N. dealbatus, N. cultriformis) and the “dorsatus group” (N. dorsatus, N. tremulus), or a mix of these. N. auletes is the largest cicada in North America. “Mega” is the Greek word for “very large” or “great”. Word is that Kathy Hill and David Marshall also planned on describing a Megatibicen genus at one point, as well.

Whenever cicada names change it causes feelings of bemusement, discontentment and discomfort amongst some cicada researches and fans. I know I don’t like it because I have to update the names of cicadas in 100’s of places on this website ;). Some folks simply disagree with the folks writing the paper. Some people prefer former names because they sound nicer (e.g. N. chloromerus vs N. tibicen tibicen). Some people simply do not like change.

Related: Here’s my article on when Neotibicen & Hadoa were established from Tibicen.

1 Hill, et al. Molecular phylogenetics, diversification, and systematics of Tibicen Latreille 1825 and allied cicadas of the tribe Cryptotympanini, with three new genera and emphasis on species from the USA and Canada (Hemiptera: Auchenorrhyncha: Cicadidae) 2015, Zootaxa 3985 (2): 219–251.

April 26, 2015

One new genus, and 15 new species of cicada in Argentina

Filed under: Allen F. Sanborn,Argentina,Maxine E. Heath — Dan @ 6:21 am

Allen F. Sanborn & Maxine S. Heath published a new paper about cicadas titled The cicadas of Argentina with new records, a new genus and fifteen new species (Hemiptera: Cicadoidea: Cicadidae) in Zootaxa Vol 3883, No 1, in November of 2014. Website for the document.

The abstract of the paper reveals some exciting discoveries:

  1. 108 species belonging to 37 genera, eight tribes, and three subfamilies of cicadas are represented in the Argentine cicada fauna.
  2. The new genus is Torresia Sanborn & Heath gen. n.
  3. New species:
    1. Adusella signata Haupt, 1918 rev. stat.
    2. Alarcta micromacula Sanborn & Heath sp. n.
    3. Chonosia longiopercula Sanborn & Heath sp. n.
    4. Chonosia septentrionala Sanborn & Heath sp. n.
    5. Dorisiana noriegai Sanborn & Heath sp. n.
    6. Fidicinoides ferruginosa Sanborn & Heath sp. n.
    7. Guyalna platyrhina Sanborn & Heath sp. n.
    8. Herrera humilastrata Sanborn & Heath sp. n.
    9. Herrera umbraphila Sanborn & Heath sp. n.
    10. Parnisa lineaviridia Sanborn & Heath sp. n.
    11. Parnisa viridis Sanborn & Heath sp. n.
    12. Prasinosoma medialinea Sanborn & Heath sp. n.
    13. Proarna alalonga Sanborn & Heath sp. n.
    14. Proarna parva Sanborn & Heath sp. n.
    15. Torresia lariojaensis Sanborn & Heath sp. n.
    16. Torresia sanjuanensis Sanborn & Heath sp. n.
  4. The document is 94 pages long.

March 25, 2014

Catalogue of the Cicadoidea by Allen F Sanborn

Filed under: Allen F. Sanborn,Books — Dan @ 7:17 pm

The Catalogue of the Cicadoidea (Hemiptera: Auchenorrhyncha) by Allen F Sanborn weighs about six pounds. It’s also one of my favorite cicada books, and it usually can be found on my desk. I use it mostly to verify the names of cicadas.

Catalogue of the Cicadoidea by Allen F Sanborn

Here’s a description from the publisher:

This is the third in a series of catalogs and bibliographies of the Cicadoidea covering 1981-2010. The work summarizes the cicada literature, providing a means for easy access to information previously published on a particular species or to allow researchers the ability to locate similar work that has been published on other species. A total of 2,591 references are included in the bibliography. The book is a source of biological and systematic information that could be used by zoologists, entomologists, individuals interested in crop protection, and students studying entomology as well as anyone interested in cicadas or who require specific information on the insects. Each genus/species is identified with the reference, the page number, any figures (if applicable), the topics covered by the reference, any synonymies, and any biogeographic information mentioned for the species in the individual reference. An added benefit to the catalog is that it is the first complete species list for the Cicadoidea, including all synonymies and new combinations through 2012.

Over 3,390 varieties of cicadas (yeah, I manually counted the species).

April 9, 2013

Biogeography of the Cicadas (Hemiptera: Cicadidae) of North America, North of Mexico

Download the PDF here: www.cicadamania.com/downloads/diversity-05-00166.pdf.

We are excited to announce the availability of a document by Allen F. Sanborn and Polly K. Phillips titled Biogeography of the Cicadas (Hemiptera: Cicadidae) of North America, North of Mexico. This document features distribution maps for North American cicada species! This document is an excellent companion to The Cicadas (Hemiptera: Cicadoidea: Cicadidae) of North America North of Mexico by Allen F. Sanborn and Maxine S. Heath (link to that book).

Abstract: We describe and illustrate the biogeography of the cicadas inhabiting continental North America, north of Mexico. Species distributions were determined through our collecting efforts as well as label data from more than 110 institutional collections. The status of subspecies is discussed with respect to their distributions. As we have shown over limited geographic areas, the distribution of individual species is related to the habitat in which they are found. We discuss the biogeography of the genera with respect to their phylogenetic relationships. California is the state with the greatest alpha diversity (89 species, 46.6% of taxa) and unique species (35 species, 18.3% of taxa). Texas, Arizona, Colorado and Utah are the states with the next greatest alpha diversity with Texas, Arizona and Utah being next for unique species diversity. Maine, New Hampshire and Rhode Island are the states with the least amount of cicada diversity. Diversity is greatest in states and areas where there is a diversity of plant communities and habitats within these communities. Mountainous terrain also coincides with increases in diversity. Several regions of the focus area require additional collection efforts to fill in the distributions of several species.
Keywords: cicada; distribution; Diceroprocta; Tibicen; Okanagana; Okanagodes; Cacama; Magicicada; Platypedia; Cicadetta

An example of a map from the document:

Example Map

December 9, 2012

Two new cicada publications worth reading

Filed under: Allen F. Sanborn,Canada,United States — Dan @ 1:46 pm

Two relatively new cicada publications that should be worth reading:

1) Avian Predation Pressure as a Potential Driver of Periodical Cicada Cycle Length by Walter D. Koenig and Andrew M. Liebhold, The American Naturalist. This is a newly electronically published paper about what drives the long, prime-numbered lifecycle of Magicicada periodical cicadas.

Abstract:

The extraordinarily long life cycles, synchronous emergences at 13- or 17-year intervals, and complex geographic distribution of periodical cicadas (Magicicada spp.) in eastern North America are a long-standing evolutionary enigma. Although a variety of factors, including satiation of aboveground predators and avoidance of interbrood hybridization, have been hypothesized to shape the evolution of this system, no empirical support for these mechanisms has previously been reported, beyond the observation that bird predation can extirpate small, experimentally mistimed emergences. Here we show that periodical cicada emergences appear to set populations of potential avian predators on numerical trajectories that result in significantly lower potential predation pressure during the subsequent emergence. This result provides new support for the importance of predators in shaping periodical cicada life history, offers an ecological rationale for why emergences are synchronized at the observed multiyear intervals, and may explain some of the developmental plasticity observed in these unique insects.

Order it from JSTOR.

2) The Cicadas (Hemiptera: Cicadoidea: Cicadidae) of N. America North of Mexico by Allen F. Sanborn and Maxine S. Heath. 227 pages.

A comprehensive review of the North Amerian cicada fauna that provides information on synonymies, type localities, and type material. There are 170 species and 21 subspecies found in continental N. America north of Mexico. The book has 211 figures with each species photographed in color.

Buy it from the Entomological Society of America website. I’ve already ordered mine.

I can’t wait for Sanborn’s book on Central and South America (hopefully, that will arrive within the next few years).

Update:

I’ve received Allen F. Sanborn and Maxine S. Heath’s book. It’s focus is on identifying all species of cicada fauna in North America, north of Mexico, as the title says. It also identifies species that were reported to exist in this location, but do not. The book provides maps and common attributes of each genus of cicada, and then for each species it provides photos of the holotype (and the location of the holotype), as well as a history of its taxonomy.

November 7, 2012

How to learn more about cicadas, by searching for cicada researchers

Filed under: Allen F. Sanborn,FAQs,Zammara — Dan @ 10:58 pm

Just about anything and everything ends up on the web these days, including research papers written by cicada researchers. Many of these papers are easily downloaded from the web, and once downloaded you can read them and expand your knowledge about cicadas.

This weekend I was looking for information about cicadas from Central and South America (the Neotropic ecozone). Allen F. Sanborn, Ph.D is well known for his research of cicadas of that region, so I searched for some of his research papers. Google will retrieve all PDF (Adobe Acrobat) files that contain the word cicada and the name Allen F. Sanborn, when you search for “Allen F. Sanborn cicada ext:pdf” (remove the quotes when you search).

Some interesting papers I found include:

Checklist of the cicadas (Insecta: Hemiptera: Cicadidae) of Paraguay including new records for six species (Sanborn, Allen F., 2011). This paper includes a long list of cicada species, which I used to look for images of cicadas on Flickr.com.

The new records increase the known cicada diversity 37.5% bringing the total number of cicada species reported in Paraguay to 22 species in 12 genera representing five tribes and three subfamilies of the family Cicadidae. There are currently no known endemic species.

Two New Zammara Species from South America (Hemiptera: Cicadomorpha: Cicadidae) (Sanborn, Allen F., Florida Entomologist 87(3),2004). This paper includes many photographs, which make cicada identification easy.

ABSTRACT
Two new members of the widespread Neotropical genus Zammara Amyot & Serville, Zammara
olivacea n.sp. from Columbia and Zammara medialinea n.sp. from Venezuela are described.

Key Words: new species, taxonomy, cicada, Zammara, Columbia, Venezuela.

New Records for the Cicada Fauna from Four Central American Countries (HEMIPTERA: CICADOIDEA: CICADIDAE (Sanborn, Allen F.; Florida Entomologist 89(1), 2006). This article features a map with cicada species names.

ABSTRACT
Analysis of museum specimens has added to the cicada fauna of Belize, El Salvador, Guate- mala, and Honduras. Information on the cicada fauna reported in the literature as well as the first records of cicada species to the fauna are reported here to provide a more accurate un- derstanding of cicada diversity in each country and the region. The new records represent an increase of 75, 14, 110, and 320%, respectively, to the cicada faunal diversity of each country.
Key Words: cicadas, biodiversity, Central America

If you use my Google formula, you can find these papers too.

The The Current Status of Cicada Taxonomy on a Region-by-Region Basis page on Cicada Central is a good resource for learning about other cicada researchers.