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October 28, 2023

Cicada research published in 2023

Filed under: Papers and Documents — Dan @ 8:13 pm

Cicada research published in 2023… this is a work in progress. 🙂


Sanborn, A, (2023). Two new species and two new records for cicadas (Hemiptera: Cicadidae) from French Guiana, with an updated faunal list of French Guiana and the first synoptic list for Guyana. Zootaxa. Vol. 5368 No. 1: 7 Nov. 2023. 1-74. DOI: 10.11646/zootaxa.5368.1.1

Stukel, M., Porczak, A.E., Gordon, E.R.L., Vailionis, J., Haji, D., Buckley, T.R. et al. (2023) Phylogenomics improves the phylogenetic resolution and provides strong evidence of mito-nuclear discordance in two genera of a New Zealand cicada (Hemiptera: Cicadidae) species radiation. Systematic Entomology, 1–21. Available from:


Hepler, J., Cooper, W., Cullum, J., Dardick, C., Dardick, L., Nixon, L., Pouchnik, D., Raupp, M., Shrewsbury, P., Leskey, R. (2023). Do adult Magicicada (Hemiptera: Cicadidae) feed? Historical perspectives and evidence from molecular gut content analysis. Journal of Insect Science, Volume 23, Issue 5, September 2023, 13,

Kaplan, Mehmet. (2023). Determination of Bioecology, Infestation Rate and Distribution of the Harmful Grapevine Cicada Klapperichicen viridissima (Walker) (Hemiptera: Cicadidae) in Vineyard Areas of Mardin Province in Türkiye. Erwerbs-Obstbau. 1-5. 10.1007/s10341-023-00961-1.


Cole, J., Chatfield-Taylor, W., Smeds, E., Cooley, J., Gonzalez, V., Wong, C. (2023). Phylogeny of North America’s largest cicada radiation redefines Tibicinoides and Okanagana (Hemiptera: Auchenorrhyncha: Cicadidae: Tibicininae). Zootaxa. Vol. 5346 No. 5: 21 Sept. 2023. DOI:


Wang, J., Jiang, J., Wei, C. (2023). A new genus, Duffelsa gen. n., with descriptions of three new species and one new combination (Hemiptera: Cicadidae). Zootaxa. Vol. 5323 No. 3: 2 Aug. 2023. DOI:

Puissant, S., Gurcel, K. (2023). The genus Tibicina Kolenati, 1857 in Morocco (Hemiptera: Cicadidae: Tibicininae): taxonomic assessment from integrative research. Zootaxa. Vol. 5330 No. 4: 17 Aug. 2023. DOI:

Diaz, A., Yaritza, L., Maes, J. (2023) Zammara calochroma (Homoptera: Cicadidae) registro nuevo para la fauna de Nicaragua. DOI:10.5281/zenodo.8233967


Sanborn, A. (2023). Resolving taxonomic issues of cicadas (Hemiptera: Cicadidae) including new combinations, new synonymies, and revised status, with updates on the diversity of the Brazilian cicada fauna and new records for four South American countries. Zootaxa. VOL. 5318 NO. 3: 20 JUL. 2023. 339-362. DOI: 10.11646/ZOOTAXA.5318.3.2


Yamna, Ouguas & Mohamed, El & Ahmed, Nani & Abdelilah, Chakhmani & Abdellah, Bouchtalla & Lhomme, Patrick. (2023). Biogeographical characterization of Psalmocharias plagifera (Schumacher, 1922) (Hemiptera: Cicadidae) mass emergence in southwest Morocco. 01-19. 10.34874/IMIST.PRSM/afrimed-i139.41148.


Sanborn, A. (2023). The taxonomic position of the cicada genus Hea Distant, 1906 (Hemiptera: Cicadidae). Zootaxa. Vol. 5254 No. 3: 13 Mar. 2023. DOI:

Lee, Y., Marshall, A., Mohagan, D., Hill, K., Mohagan, D. (2023). Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species. Journal of Natural History. 57(1-4):193-242. DOI:10.1080/00222933.2023.2171820

Lampert, E., Perez, G., Alejo, D., Jones, S., Ignatius, A. (2023). The 2021 Emergence of Brood X Periodical Cicadas Magicicada spp. (Hemiptera: Cicadidae) in Georgia, United States of America. Environmental Entomology, 52(2):270-278 (2023).


Wang, C. (2023). A new species of Hea Distant, 1906 from China, with annotated catalogue for the genus (Hemiptera: Cicadidae: Cicadettinae). Zootaxa. Vol. 5231 No. 5: 2 Feb. 2023. DOI:

Dmitriev, D., Sanborn, A. (2023). Priority and the names of two cicada tribes (Hemiptera: Cicadidae: Cicadinae: Tacuini and Tettigomyiinae: Anopercalnini). Zootaxa. Vol. 5239 No. 1: 8 Feb. 2023. DOI:


Chantarachit, R., Srikosamatara, S. (2023). Emergence Pattern of Dundubia nagarasingna (Hemiptera: Cicadidae) in a Bangkok Urban Garden. J. of Agricultural and Urban Entomology, 38(1):41-51 (2023).


Research from other years: 2022, 2021, 2020, 2019, 2018, 2017.

March 27, 2022

Cicada research published in 2022

Filed under: Papers and Documents — Dan @ 6:26 pm

This is a list of cicada-related research published so far in 2022. 46!

November 2022

  1. Haji, D., Vailionis, J., Stukel, M. et al. Lack of host phylogenetic structure in the gut bacterial communities of New Zealand cicadas and their interspecific hybrids. Sci Rep 12, 20559 (2022).

October 2022

  1. Allen M. Young, Aiden S. Mahoney, and Jason Canfield “A Multi-year Adult Emergence Study of the Cicada Neotibicen canicularis (Harris) (Hemiptera: Cicadidae) in Wisconsin,” The American Midland Naturalist 188(2), 250-258, (20 October 2022).
  2. Hasan, Cecilia & Sutton, Reagan & Replogle, Jessica. (2022). The Presence of Wolbachia in Brood X Cicadas.
  3. Brumfield, K.D., Raupp, M.J., Haji, D. et al. Gut microbiome insights from 16S rRNA analysis of 17-year periodical cicadas (Hemiptera: Magicicada spp.) Broods II, VI, and X. Sci Rep 12, 16967 (2022).
  4. Costa, G. J., Nunes, V. L., Marabuto, E., Mendes, R., Silva, D. N., Pons, P., Bas, J. M., Hertach, T., Paulo, O. S., & Simões, P. C. (2023). The effect of the Messinian salinity crisis on the early diversification of the Tettigettalna cicadas. Zoologica Scripta, 52, 100– 116.
  5. Allison M Roth, Sarah M Kent, Elizabeth A Hobson, Gene Kritsky, Shinichi Nakagawa, Personality-mediated speed-accuracy tradeoffs in mating in a 17-year periodical cicada, Behavioral Ecology, Volume 33, Issue 6, November/December 2022, Pages 1141–1152.
  6. Hoa Quynh Nguyen, Erick Kim, Yoonhyuk Bae, Soyeon Chae, Seongmin Ji, Jiman Heo, Sungsik Kong, Thoa Kim Nguyen, Thai Hong Pham, Yikweon Jang, An effective method for accurate nymphal-stage delimitation of the cicada Hyalessa fuscata, Journal of Asia-Pacific Entomology, Volume 25, Issue 3, 2022, 101952,

September 2022

  1. Bergh, James & Nita, M. & Dyer, J.E. & Brandt, S.N. & Cullum, John & Nixon, Laura & Leskey, Tracy. (2022). Spatial distribution of 17-year periodical cicada (Hemiptera: Cicadidae) exuviae and oviposition injury in Mid-Atlantic, USA Apple orchards and implications for management. Crop Protection. 162. 106095. 10.1016/j.cropro.2022.106095.
  2. Diler Haji, Jason Vailionis, Mark Stukel et al. Correlates of host-associated bacterial diversity in New Zealand cicadas and hybrids, 22 September 2022, PREPRINT (Version 1) available at Research Square []
  3. Kriesberg, Caleb. (2022). Protandrous Arrival in a Population of the Periodical Cicada Magicicada septendecim (Linnaeus) (Hemiptera: Cicadidae) in Montgomery County, Maryland: Addendum to METHODS and RESULTS Sections.
  4. J.C. Bergh, M. Nita, J.E. Dyer, S.N. Brandt, J.P. Cullum, L.J. Nixon, T.C. Leskey, Spatial distribution of 17-year periodical cicada (Hemiptera: Cicadidae) exuviae and oviposition injury in Mid-Atlantic, USA Apple orchards and implications for management, Crop Protection, Volume 162, 2022, 106095.
  5. Max Moulds, Michael Frese & M. R. McCurry (2022) New cicada fossils from Australia (Hemiptera: Cicadoidea: Cicadidae) with remarkably detailed wing surface nanostructure, Alcheringa: An Australasian Journal of Palaeontology,
  6. Kriesberg, Caleb. (2022). Emergence Patterns and Species Distribution of the Brood X 17-Year Periodical Cicada, Magicicada, Davis (Hemiptera: Cicadidae), near Downtown Silver Spring, Montgomery County, Maryland, 2021. 8. 2.
  7. Gurcel, Kevin & Stéphane, Puissant. (2022). Une Cigale dans le blanc des yeux : synopsis d’une aberration chromatique très rare chez Cicadetta montana (Scopoli, 1772) (Hemiptera, Cicadidae). Bulletin de la Société entomologique de France. 127. 259-271.

August 2022

  1. Moulds, Max & MARSHALL, DAVID. (2022). New genera and new species of Western Australian cicadas (Hemiptera: Cicadidae). Zootaxa. 5174. 451-507. 10.11646/zootaxa.5174.5.1.
  2. Takahiro Ishimaru, Ikkyu Aihara. (2022). Temporal structure of two call types produced by competing for male cicadas. bioRXiv. (preprint).
  3. Sota, T. (2022). Life-cycle control of 13- and 17-year periodical cicadas: A hypothesis and its implication in the evolutionary process. Ecological Research, 37( 6), 686– 700.
  4. Cassandra L. Ettinger, Brian Lovett, Matt T. Kasson, Jason E. Stajich. (2022). Metagenome-Assembled Genomes of Bacteria Associated with Massospora cicadina Fungal Plugs from Infected Brood VIII Periodical Cicadas. Microbiology Resource Announcements. e00413-22. 11. 10.
  5. Jason E. Stajich, Brian Lovett, Cassandra L. Ettinger, Derreck A. Carter-House, Tania Kurbessoian, Matt T. Kasson. (2022). An Improved 1.5-Gigabase Draft Assembly of Massospora cicadina (Zoopagomycota), an Obligate Fungal Parasite of 13- and 17-Year Cicadas. Microbiology Resource Announcements. e00367-22. 11. 10.

July 2022

June 2022

  1. Giroux, Marjolaine & Legault, Audrey & Bede, J.C.. (2022). Flesh flies (Diptera: Sarcophagidae) attracted to dog-day cicada (Neotibicen canicularis (Harris) Hemiptera: Cicadidae) carcasses in Québec, Canada. 2022.
  2. Hong, Jung-Hee & Lee, Young-Cheol. (2022). Anti-Inflammatory Effects of Cicadidae Periostracum Extract and Oleic Acid through Inhibiting Inflammatory Chemokines Using PCR Arrays in LPS-Induced Lung inflammation In Vitro. Life. 12. 857. 10.3390/life12060857.
  3. Lalremsanga, H.T. & Muansanga, Lal & Malsawmdawngliana, Fanai & Khawlhring, Marova. (2022). A report on the occurrence of the cicada Callogaeana festiva (Fabricius, 1803) (Insecta: Cicadidae) from Mizoram, India. Journal of Threatened Taxa. 14. 21321-21323. 10.11609/jott.7550.14.6.21321-21323.
  4. Li, Qian & Ji, Aihong & Shen, Huan & Han, Qingfei & Qin, Guodong. (2022). The forewing of a black cicada Cryptotympana atrata (Hemiptera, Homoptera: Cicadidae): Microscopic structures and mechanical properties. Microscopy Research and Technique. 85. 10.1002/jemt.24173.
  5. Luu, Hoang & Pham, Thai & Bui, Thu. (2022). Research on assessment of the diversity and similarity of cicadas (Hemiptera: Cicadidae) in the Northwest region. Ministry of Science and Technology, Vietnam. 64. 27-31. 10.31276/VJST.64(1).27-31.
  6. Luu, Hoang & Pham, Thai & Bui, Thu. (2022). The composition and distribution of the Cicadidae family (Hemiptera: Auchenorrhyncha) from Northwestern in Vietnam. Ministry of Science and Technology, Vietnam. 64. 24-27. 10.31276/VJST.64(6).24-27.
  7. Mora-Rubio, Carlos. (2022). Contributions to the distribution of the Iberian endemism Hilaphura varipes (Hemiptera, Cicadidae). Boletin – Asociacion Espanola de Entomologia. 46. 77-82.
  8. Owen, Christopher & Marshall, David & Wade, Elizabeth & Meister, Russ & Goemans, Geert & Kunte, Krushnamegh & Moulds, Max & Hill, Kathy & Villet, Martin & Pham, Thai & Kortyna, Michelle & Lemmon, Emily & Lemmon, Alan & Simon, Chris. (2022). Detecting and Removing Sample Contamination in Phylogenomic Data: An Example and its Implications for Cicadidae Phylogeny (Insecta: Hemiptera). Systematic Biology. 10.1093/sysbio/syac043.
  9. Wang Cheng-Bin, ??? & Liu, Peng-Yu. (2022). A new species of Polyneura Westwood, 1842 from Yunnan, China (Hemiptera, Cicadidae, Cicadinae). Biodiversity Data Journal. 10. 84554. 10.3897/BDJ.10.e84554. Image.

May 2022

  1. Brown, Daniel & Kotsani, Natalia. (2022). A Stochastic Modeling of the Cicada Chorus.
  2. Figueroa-Rodríguez, Rosa & Gálvez-Marroquín, Luis & Martinez, Misael & Cruz López, Jesús Alberto & Ariza-Flores, Rafael & Alonso, Moises & Sánchez-García, José & García-Mayoral, Luis. (2022). Quantification of direct and indirect damage caused by Diceroprocta bulgara (Distant) (Hemiptera: Cicadidae) in lime. Agro Productividad. 10.32854/agrop.v15i4.2054.
  3. Heath, James & Heath, Maxine & Sanborn, Allen. (2022). Cold cicadas and hot rocks: Thermal responses and thermoregulation in some New Zealand cicadas (Insecta: Hemiptera: Cicadidae: Cicadettinae: Cicadettini). Journal of Thermal Biology. 107. 103273. 10.1016/j.jtherbio.2022.103273.
  4. Lee, Young June. (2022). A new genus and species of the subtribe Leptopsaltriina (Hemiptera: Cicadidae: Leptopsaltriini) from Sabah, Malaysia. Journal of Asia-Pacific Biodiversity. 15. 10.1016/j.japb.2022.05.006.
  5. Popple, Lindsay & Emery, David. (2022). Five new species of Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae) from south-eastern Australia. Zootaxa. 5141. 401-441. 10.11646/zootaxa.5141.5.1.
  6. Setälä, Heikki, Szlavecz, Katalin, Pullen, Jamie D., Parker, John D., Huang, Yumei, Chang, Chih-Han. 2022. Acute Resource Pulses from Periodical Cicadas Propagate to Belowground Food Webs but Do Not Affect Tree Performance. Ecology e3773.

April 2022

  1. Marshall, David C. On the spelling of the name of Cassin’s 17-Year Cicada, Magicicada cassini (Fisher, 1852) (Hemiptera: Cicadidae). 2022. Zootaxa 5125 (2): 241–245. 10.11646/zootaxa.5125.2.8
  2. Moulds, Max & Marshall, David & Hutchinson, Paul. (2022). Pericallea katherina, a new cicada genus and species from Western Australia (Hemiptera: Cicadidae: Cicadettini). Australian Entomologist. 49. 1-14.
  3. Shi, Peijian & Jiao, Yabing & Niklas, Karl & Li, Yirong & Guo, Xuchen & Yu, Kexin & Chen, Long & Hurd, Lawrence. (2022). Sexual Dimorphism in Body Size and Wing Loading for Three Cicada Species. Annals of the Entomological Society of America.

March 2022

  1. Belenguier, Luc. (2022). Tettigettalna argentata (Olivier 1790) (Hemiptera: Cicadidae), une nouvelle cigale pour le département de l’Allier. BIOM – Revue scientifique pour la biodiversité du Massif central. 3.
  2. Emery, David & Emery, Nathan & Hutchinson, Paul & Ong, Simon. (2022). Two new species of Tryella Moulds, 2003 (Hemiptera: Cicadidae) from Western Australia and Northern Territory. Australian Entomologist. 49. 23-42.
  3. Liang, Shih-Hsiung & Lee, Lin-Lee & Shieh, Bao-Sen. (2022). Female preference for song frequency in the cicada Mogannia formosana Matsumura (Hemiptera: Cicadidae). Behavioural Processes. 197. 104626.
  4. Maes, Jean. (2022). Zammara smaragdula (Homoptera: Cicadidae) reporte nuevo para la fauna de Nicaragua 1. REVISTA NICARAGUENSE DE ENTOMOLOGIA, 268.. 267. 1-22. 10.5281/zenodo.6554509.
  5. Yen, Luu & Pham, Thai & Constant, Jérôme. (2022). A NEW SPECIES OF Platylomia Stål, 1870 (Hemiptera: Cicadidae) FROM VIETNAM, WITH A KEY TO SPECIES. ACADEMIA JOURNAL OF BIOLOGY. 44. 23-31. 10.15625/2615-9023/16757.

February 2022

  1. ?, ??. (2022). Enrichment and Screening of Several Micronutrients in Cicada Flower Fruiting Body. Hans Journal of Food and Nutrition Science. 11. 21-26.
  2. Perkovich, C., & Ward, D. (2022). Changes in white oak (Quercus alba) phytochemistry in response to periodical cicadas: Before, during, and after an emergence. Ecology and Evolution, 12, e8839.
  3. Shi, Cuie & Song, Wenlong & Gao, Jian & Yan, Shoubao & Guo, Chen & Zhang, Tengfei. (2022). Enhanced production of cordycepic acid from Cordyceps cicadae isolated from a wild environment. Brazilian Journal of Microbiology.

January 2022

  1. Hoang Yen Luu, Hong Thai Pham, Thu Quynh Bui. Research on assessment of the diversity and similarity of cicadas (Hemiptera: Cicadidae) in the Northwest region. Vietnam Forest Museum, Forestry Inventory and Planning Institute. Vietnam National Museum of Nature, VAST3. Graduate University of Science and Technology, VAST Received 8 November 2021; accepted 14 December 2021.

November 24, 2021

Cicada Research Published in 2021

Filed under: Papers and Documents — Dan @ 10:36 pm

This is a list of cicada-related research published in 2021.

More than 60 so far (12/24/2021). Email with any more.


  1. Dandan, Wang & Huang, Zhi & Billen, Johan & Zhang, Guoyun & He, Hong & Wei, Cong. (2021). Complex co-evolutionary relationships between cicadas and their symbionts. Environmental Microbiology. 10.1111/1462-2920.15829. Link.
  2. Kritsky, Gene. (2021). One for the Books: The 2021 Emergence of the Periodical Cicada Brood X. American Entomologist. 67. 40-46. 10.1093/ae/tmab059. Link.
  3. Tomita, Kanji & Makoto, Kobayashi. (2021). Development of experimental mesocosms for cicada nymphs Graptopsaltria nigrofuscata: methodology and research recommendations. 93. 207-212. 10.25674/so93iss3id171. Link.
  4. Moulds, Max. (2021). A revision of the endemic Australian cicada genus Jassopsaltria Ashton, 1914 (Cicadidae: Cicadinae: Jassopsaltriini). Australian Entomologist. 48. 217-240. Link.
  5. Moriyama, Minoru & Yasuyama, Kouji & Numata, Hideharu. (2021). The formation of a hatching line in the serosal cuticle confers multifaceted adaptive functions on the eggshell of a cicada. Zoological Letters. 7. 10.1186/s40851-021-00178-8. Link.
  6. Tomita, Kanji. (2021). Camera traps reveal interspecific differences in the diel and seasonal patterns of cicada nymph predation. The Science of Nature. 108. 10.1007/s00114-021-01762-w. Link.
  7. Tian, Lejie & Liu, Hui & Zhang, Beiyi & Liu, Yi & Lv, Siye & Pang, Lingyan & Li, Junqi. (2021). Ni and CeO 2 Nanoparticles Anchored on Cicada-Wing-like Nitrogen-Doped Porous Carbon as Bifunctional Catalysts for Water Splitting. ACS Applied Nano Materials. 10.1021/acsanm.1c03850. Link.
  8. Sanborn, Allen & Cole, Jeffrey & Stukel, Mark & Lukasik, Piotr & Veloso, Claudio & Gonzalez, Valorie & Karkar, Jessica & Simon, Chris. (2021). Thirteen new species of Chilecicada Sanborn, 2014 (Hemiptera: Auchenorrhyncha: Cicadidae: Tibicininae) expand the highly endemic cicada fauna of Chile. Zootaxa. 5078. 1-70. 10.11646/zootaxa.5078.1.1. Link.


  1. Kojevnikova, A.. (2021). The Study of Cicadas in the Conditions of the Fergana Valley of Uzbekistan. International Journal of Multidisciplinary Research and Analysis. 04. 10.47191/ijmra/v4-i11-38. Link.
  2. Huang, Ailin & Wu, Tao & Wu, Xiuyun & Zhang, Biao & Shen, Yuanyuan & Wang, Suying & Song, Wenjun & Ruan, Haihua. (2021). Analysis of Internal and External Microorganism Community of Wild Cicada Flowers and Identification of the Predominant Cordyceps cicadae Fungus. Frontiers in microbiology. 12. 752791. 10.3389/fmicb.2021.752791. Link.
  3. Mozaffarian, Fariba. (2021). Cicadas and hoppers as pests of field crops in Iran (Hemiptera: Auchenorrhyncha). Link.
  4. Mukaimine, Wataru & Kawatsu, Kazutaka & Toquenaga, Yukihiko. (2021). Digging out intersexual and meteorological effects on cicada emergence using 10-year citizen monitoring. Ecological Entomology. 10.1111/een.13109. Link.
  5. Poerio, Aurelia & Girardet, Thomas & Petit, Chloé & Fleutot, Solenne & Jehl, Jean-Philippe & Arab-Tehrany, Elmira & Mano, João F. & Cleymand, Franck. (2021). Comparison of the Physicochemical Properties of Chitin Extracted from Cicada orni Sloughs Harvested in Three Different Years and Characterization of the Resulting Chitosan. Applied Sciences. 11. 11278. 10.3390/app112311278. Link.


  1. Lee, Young & Mohagan, Alma. (2021). A new species of Oncotympana Stål, 1870 (Hemiptera: Cicadidae: Oncotympanini) from Negros, Philippines. Journal of Asia-Pacific Biodiversity. Link.
  2. Simon, Chris & Cooley, John & Karban, Richard & Sota, Teiji. (2021). Advances in the Evolution and Ecology of 13- and 17-Year Periodical Cicadas. Annual review of entomology. 67. 10.1146/annurev-ento-072121-061108. Link.
  3. Nagata, Nobuaki & Toda, Mamoru & Ohbayashi, Takashi & Hayashi, Masami & Sota, Teiji. (2021). Phylogeography of cicadas on continental and oceanic islands in the northwestern Pacific region. Journal of Biogeography. 48. 10.1111/jbi.14262. Link.
  4. Umam, Rofiqul & Kinarya P, Endah & Yakin, Khusnul & Iqbal, Mochamad & Siregar, Rahmat nawi & Husein, Irzaman. (2021). Intensity level as sustainable energy: Analysis of the conversion of energy stored on cicadas sound waves. Jurnal Ilmiah Pendidikan Fisika Al-Biruni. 10. 35-44. 10.24042/jipfalbiruni.v10i2.9798. Link.


  1. Moulds, Max & Marshall, D & Popple, Lindsay. (2021). Kimberpsaltriini, a new tribe for a new Australian cicada allied to Talcopsaltria Moulds (Hemiptera: Cicadoidea: Cicadidae). Australian Entomologist. 48. 149-160. Link.
  2. Zhang, Lijia & Wang, Siyue & Billen, Johan & Wei, Cong. (2021). Morphology and ultrastructure of the epithelial femoral gland in cicadas (Hemiptera: Cicadidae). Arthropod Structure & Development. 64. 101086. 10.1016/j.asd.2021.101086. Link.
  3. Hajong, Sudhanya & Limatemjen,. (2021). Platylomia kohimaensis n. sp.—a new cicada species (Hemiptera: Cicadidae) from the Naga Hills in the Eastern Himalayas. Zootaxa. 5047. 081-091. 10.11646/zootaxa.5047.1.7. Link.
  4. Kalesh, S.. (2021). A new species of Pomponia Stål, 1866 (Hemiptera: Cicadidae) from the Western Ghats, with notes on the status of P. linearis (Walker, 1850) from southern India. Zootaxa. 5040. 388-403. 10.11646/zootaxa.5040.3.4. Link.
  5. Bator, John & Marshall, David & Hill, Kathy & Cooley, John & Leston, Adam & Simon, Chris. (2021). Phylogeography of the endemic red-tailed cicadas of New Zealand (Hemiptera: Cicadidae: Rhodopsalta ), and molecular, morphological and bioacoustical confirmation of the existence of Hudson’s Rhodopsalta microdora. Zoological Journal of the Linnean Society. 10.1093/zoolinnean/zlab065. Link.
  6. Zhang, Lijia & Wang, Siyue & Billen, Johan & Wei, Cong. (2021). Morphology and ultrastructure of the epithelial femoral gland in cicadas (Hemiptera: Cicadidae). Arthropod Structure & Development. 64. 101086. 10.1016/j.asd.2021.101086. Link.
  7. Nguyen, Hoa & Kim, Yuseob & Yikweon, Jang. (2021). De Novo Transcriptome Analysis Reveals Potential Thermal Adaptation Mechanisms in the Cicada Hyalessa fuscata. Animals. 11. 2785. 10.3390/ani11102785. Link.
  8. Qian, Yi & Sun, Xin & Wang, Xin & Yang, Xin & Fan, Mengyao & Zhong, Jiao & Pei, Zejun & Guo, Junping. (2021). Mechanism of Cordyceps Cicadae in Treating Diabetic Nephropathy Based on Network Pharmacology and Molecular Docking Analysis. Journal of Diabetes Research. 2021. 1-10. 10.1155/2021/5477941. Link.


  1. Villet, Martin & Edwards, Shelley. (2021). The cicada genus Tugelana Distant, 1912 (Hemiptera, Cicadidae): phylogenetic position and conservation status. African Invertebrates. 62. 399-410. 10.3897/afrinvertebr.62.66891. Link.
  2. Haji, Diler & Vailionis, Jason & Stukel, Mark & Gordon, Eric & Lemmon, Emily & Lemmon, Alan & McCutcheon, John & Simon, Chris. (2021). Host-associated microbial diversity in New Zealand cicadas uncovers elevational structure and replacement of obligate bacterial endosymbionts by Ophiocordyceps fungal pathogens. 10.1101/2021.08.24.457591. Link.


  1. Zhao, Chenchen. (2021). High-throughput sequencing yields a complete mitochondrial genome of the Cryptotympana atrata (Insecta: Hemiptera: Cicadidae). Mitochondrial DNA Part B. 6. 1883-1885. 10.1080/23802359.2021.1934154. Link.
  2. Pham, Thai & Lee, Young June. (2021). A new genus and species of the subtribe Leptopsaltriina (Hemiptera: Cicadidae: Leptopsaltriini) from Vietnam, with a key to the genera of Leptopsaltriina. Journal of Asia-Pacific Entomology. 10.1016/j.aspen.2021.07.011. Link.
  4. Sousa P, Grosso-Silva JM, Andrade R, Chaves C, Pinto J, Paupério J, Beja P, Ferreira S (2021) The InBIO Barcoding Initiative Database: DNA barcodes of Portuguese Hemiptera 01. Biodiversity Data Journal 9: e65314. Link.
  5. Malakoff, David & Stokstad, Erik. (2021). Something is killing U.S. birds. It’s not cicadas. Science. 373. 146-146. 10.1126/science.373.6551.146. Link.
  6. Kojevnikova, Alevtina. (2021). HARMFUL CICADA GENUS EMPOASCA WALSH OF THE FERGANA VALLEY. Universum:Chemistry & biology. 84. 10.32743/UniChem.2021.84.6.11700. Link.
  7. Lao, Thuan & Trinh, Van Hanh & Vuong, Loi & Vu Tien, Luyen & Le, Thuy & Dinh, Hiep & Truong, Nguyen. (2021). Molecular record for the first authentication of Isaria cicadae from Vietnam. Open Life Sciences. 16. 711-718. 10.1515/biol-2021-0074. Link.


  1. Escola, João & Guido, Rodrigo & Silva, Ivan & Maccagnan, Douglas & Cardoso, Alexandre & Souza, Uender. (2021). Estado da arte no monitoramento acústico de Cicadidae em lavouras de café: State of the art in acoustic monitoring of Cicadidae in coffee crops. Revista Macambira. 5. e051007. 10.35642/rm.v5i1.562. Link.
  2. Emery, Nathan & Emery, David. (2021). A new cicada species, Haemopsalta eximia (Hemiptera: Cicadidae) from north-western Sydney, Australia. Australian Entomologist. 48. 97. Link.
  3. Sanborn, Allen & Allick, Earl & Apang, Sandee & Castillo, Izyanna & Cruz, Erica & Davis, Theophilus & Duncan, Cory & Fierro, Fanny & Gebaide, Marla & Luke, Abigail & Pacheco, Maria & Paz-Castillo, Daniel & Perez, Laura & Poeck, Ana & Seepersaud, Adrian & Valdes, Carolina. (2021). Flight system morphology and minimum flight temperature in North American cicadas (Insecta: Hemiptera: Cicadidae). Journal of Thermal Biology. 100. 103004. 10.1016/j.jtherbio.2021.103004. Link.
  4. Pons, Pere & Carbonell-Font, Rafael & Franch, Martí & Lay, Josep & Espejo Fraga, Daniel & Fontelles, Ferran & Funosas, David & Fusellas, Marc & Puig-Gironès, Roger & Tobella, Carles & Franch, Marc. (2021). Diversitat, distribució i fenologia de les cigales (Hemiptera: Cicadidae) a Catalunya (NE Península Iberica) – Diversity, distribution and phenology of cicadas (Hemiptera: Cicadidae) in Catalonia (NE Iberian Peninsula) – Butlletí de la Institució Catalana d’Història Natural. 85. 59. 10.2436/20.1502.01.74. Link.
  5. WANG, SIYUE & PENG, XIAODONG & Wei, Cong. (2021). A review of the cicada genus Haphsa Distant, 1905 with the description of one new species from China (Hemiptera: Cicadidae) and a phylogenetic analysis of Haphsa and its allies. Zootaxa. 4991. 523-538. 10.11646/zootaxa.4991.3.5. Link.
  6. Li J, Liu H, Wu Y, Ye L, Huang X (2021) A dataset on type specimens of hemipteran insects in China. Biodiversity Data Journal 9: e64443. Link.
  7. Crespi, Sarah. (2021). Cicada citizen science, and expanding the genetic code. Science. 10.1126/science.abj7361. Link.
  8. Tian, Juanjuan & Zhang, Cangping & Wang, XiaoMeng & Rui, Xin & Zhang, Qiuqin & Chen, Xiaohong & Dong, Mingsheng & Li, Wei. (2021). Structural characterization and immunomodulatory activity of intracellular polysaccharide from the mycelium of Paecilomyces cicadae TJJ1213. Food Research International. 147. 110515. 10.1016/j.foodres.2021.110515. Link.


  1. Dandan, Wang & Liu, Yunxiang & Yan, Su & Wei, Cong. (2021). Bacterial Communities in Bacteriomes, Ovaries and Testes of three Geographical Populations of a Sap-Feeding Insect, Platypleura kaempferi (Hemiptera: Cicadidae). Current Microbiology. 78. 10.1007/s00284-021-02435-7. Link.
  2. Cen, Qi-Wen & Wang, Zheng-Yun & Tang, Zhen-Xing & Zhang, Yu & Chen, Tao & Xue, Da-Wei & Xu, Ming-Feng & Bai, Xue-Lian & Zhou, Ting & Shi, Lu-E. (2021). Initial Purification of Antimicrobial Fermentation Metabolites from Paecilomyces cicadae and Its Antimicrobial Mechanism. LWT. 148. 111785. 10.1016/j.lwt.2021.111785. Link.
  3. Su, Qihui & Zhang, Zhicai & Liu, Xiaocui & Wang, Feng. (2021). The transcriptome analysis on urea response mechanism in the process of ergosterol synthesis by Cordyceps cicadae. Scientific Reports. 11. 10927. 10.1038/s41598-021-90377-2. Link.
  4. Zeng, Zhaoying & Mou, Dan & Luo, Li & Zhong, Wenlin & Duan, Lin & Xiao, Zou. (2021). Different Cultivation Environments Affect the Yield, Bacterial Community and Metabolites of Cordyceps cicadae. Frontiers in Microbiology. 12. 669785. 10.3389/fmicb.2021.669785. Link.


  1. EWART, A. & Moulds, Max. (2021). An impressive new species of Mugadina from Queensland with notes on a congeneric species (Cicadidae: Cicadettinae: Cicadettini). Zootaxa. 4963. 563-576. 10.11646/zootaxa.4963.3.9. Link.
  2. Cooley, John & Marshall, David & Simon, Chris. (2021). Documenting Single-Generation Range Shifts of Periodical Cicada Brood VI (Hemiptera: Cicadidae: Magicicada spp.). Annals of the Entomological Society of America. 114. 10.1093/aesa/saab007. Link.
  3. Sanborn, Allen. (2021). A new species, genus and tribe of cicada (Hemiptera: Cicadoidea: Cicadidae: Tibicininae) from Chile with a list of Chilean cicada fauna. Zootaxa. 4952. 87-100. 10.11646/zootaxa.4952.1.5. Link.
  4. Reid, Graham & McCormack, James & Habimana, Olivier & Bayer, Fabian & Goromonzi, Catherine & Casey, Eoin & Cowley, Aidan & Kelleher, Susan. (2021). Biomimetic Polymer Surfaces by High Resolution Molding of the Wings of Different Cicadas. Materials. 14. 1910. 10.3390/ma14081910. Link.


  1. WANG, JIALI & DUFFELS, JOANNES & WEI, CONG. (2021). Description of a new species of the genus Maua Distant (Hemiptera, Cicadidae) from China. Zootaxa. 4949. 582-588. 10.11646/zootaxa.4949.3.8. Link.
  2. Lee, Young June. (2021). A new species of Platypleura Amyot & Audinet-Serville (Insecta: Hemiptera: Cicadidae: Platypleurini) from Mindanao, Philippines. Journal of Asia-Pacific Biodiversity. 14. 10.1016/j.japb.2021.03.003. Link.
  3. Cole, Jeffrey. (2021). Bioacoustics of California Cacama Distant, 1904 with the first state record of C. moorei Sanborn & M. Heath, 2011 (Hemiptera: Auchenorrhyncha: Cicadidae). The Pan-Pacific Entomologist. 97. 10.3956/2021-97.1.13. Link.
  4. Pham, Thai & Lee, Young June. (2021). A new genus and species of the subtribe Aolina (Hemiptera: Cicadidae: Dundubiini) from Vietnam, with discussion on the taxonomic positions of Meimuna-like genera and species groups. Journal of Asia-Pacific Entomology. 24. 10.1016/j.aspen.2021.02.012. Link.


  1. Ruschel, Tatiana & Sanborn, Allen. (2021). Two new genera and species of Fidicinini Distant, 1905 with a re-description of Nosola Stål, 1866 (Hemiptera, Auchenorrhyncha, Cicadidae). Zootaxa. 4920. 509-527. 10.11646/zootaxa.4920.4.3. Link.
  2. Sanborn, Allen. (2021). The cicadas (Hemiptera: Cicadoidea: Cicadidae) of Madagascar including a new tribe, five new genera, twelve new species, four new species synonymies, five revised species status, ten new combinations, new tribal assignments for four genera, one new subtribe synonymy, a checklist and key to the species Zootaxa. Vol. 4937 No. 1: 24. Link.


  1. Thapa, Punam & Katila, Nikita & Choi, Hyukjae & Han, Ah-Reum & Choi, Dong-Young & Nam, Joowon. (2021). Neuroprotective Effects of N-Acetyldopamine Dimers from Cicadidae Periostracum. Link.
  2. Mahmoudikordi, Farzaneh & Balvardi, Mohammad & Akhavan, Hamid-Reza. (2021). Optimization of Ethanol-Assisted Aqueous Oil Extraction from a Cicadidae Sp. 10.1101/2021.01.24.427958. Link.
  3. Sanborn, Allen & Villet, Martin. (2021). A New Species of Buyisa Distant, 1907 (Hemiptera: Cicadidae: Cicadettinae: Cicadettini) from South Africa. Proceedings of the Entomological Society of Washington. 123. 10.4289/0013-8797.123.1.199. Link.
  4. Park, Gunhyuk & Moon, Byeong & Ryu, Seung & Kim, Wook & Lim, Hye-Sun. (2021). Cicadidae Periostracum Attenuates Atopic Dermatitis Symptoms and Pathology via the Regulation of NLRP3 Inflammasome Activation. Oxidative Medicine and Cellular Longevity. 2021. 1-16. 10.1155/2021/8878153. Link.
  5. Hill, Kathy & Marshall, David & Marathe, Kiran & Moulds, Max & Lee, Young June & Pham, Thai & Mohagan, Alma & Sarkar, Vivek & Price, Benjamin & Duffels, J. & Schouten, Marieke & Boer, Arnold & Kunte, Krushnamegh & Simon, Chris. (2021). The molecular systematics and diversification of a taxonomically unstable group of Asian cicada tribes related to Cicadini Latreille, 1802 (Hemiptera : Cicadidae). Invertebrate Systematics. 35. 10.1071/IS20079. Link.
  6. Allen F. Sanborn “A New Genus and Species of Neotropical Taphurini Distant, 1905 (Hemiptera: Cicadoidea: Cicadidae: Cicadettinae) from Brazil with a Note on the Taxonomic Position of Prosotettix Jacobi, 1907,” Proceedings of the Entomological Society of Washington, 123(1), 190-198, (29 January 2021). Link.
  7. Sarkar, Vivek & Mahapatra, Cuckoo & Mohapatra, Pratyush & Kunte, Krushnamegh. (2021). Description of three new species of the genus Mata Distant, 1906 (Hemiptera: Cicadidae: Cicadinae: Oncotympanini) with notes on their natural history from Indian state of Meghalaya, India. Zootaxa. 4908. 1-28. 10.11646/zootaxa.4908.1.1. Link.
  8. Kye, Geunho & Machta, Jonathan & Abbott, Karen & Hastings, Alan & Huffmyer, William & Ji, Fang & Liebhold, Andrew & Blackwood, Julie. (2021). Sharp boundary formation and invasion between spatially adjacent periodical cicada broods. Journal of Theoretical Biology. 515. 110600. 10.1016/j.jtbi.2021.110600. Link.

July 12, 2021

A new genus Vittagaeana and a new Cicada paper to go with it

Filed under: Gaeanini | Papers and Documents | Vittagaeana — Dan @ 12:51 pm

Recently a new paper was published with the authors Kathy B. R. Hill, David C. Marshall, Kiran Marathe, Maxwell S. Moulds, Young June Lee, Thai-Hong Pham, Alma B. Mohagan, Vivek Sarkar, Benjamin W. Price, J. P. Duffels, Marieke A. Schouten, Arnold J. de Boer, Krushnamegh Kunte and Chris Simon titled The molecular systematics and diversification of a taxonomically unstable group of Asian cicada tribes related to Cicadini Latreille, 1802 (Hemiptera: Cicadidae) in Invertebrate Systematics, 2021, 35, 570—601. Link to the paper.

Here’s the abstract:

Abstract. The cicadas (Hemiptera: Cicadidae) related to tribe Cicadini exhibit some of the most remarkable phenotypes in the family, with many genera possessing striking colour patterns and unusual morphological features. This largely Asian group of 13 tribes has proven challenging for cicada taxonomists, in part because of likely convergent evolution or losses of these phenotypes. We present the first focused molecular phylogeny of this clade, including ~60 described genera. The genetic dataset contains 839 ingroup-informative sites (out of 2575) from mitochondrial cytochrome c oxidase subunit I, nuclear elongation factor-1 a, and nuclear acetyltransferase. We use Bayesian and maximum likelihood trees to test recent changes in tribe- and subtribe-level classification, and we reconstruct ancestral character states for potentially convergent traits influencing tribe descriptions. We use fossil and molecular clock calibrations to estimate the temporal and geographic context of the radiation. The tribes Gaeanini, Leptopsaltriini, Platypleurini, Psithyristriini, and Tosenini appear polyphyletic and in need of revision, in part because of convergent evolution of opaque wings and multiple convergent gains or losses of abdominal tubercles. Kalabita Moulton, 1923 is transferred from Platypleurini to Leptopsaltriini. Vittagaeana gen. nov. is established for Vittagaeana paviei comb. nov. and Vittagaeana dives comb. nov., formerly in Tosena. Sinosenini syn. nov. is synonymised with

As part of the paper, we get a new genus, Vittagaeana, made up of two cicadas formerly belonging to the genus Tosena. They were likely placed under Tosena because of the white line on their forewings, but after a review of genetics and other features, they have been given their own genus. They are closely related to the genera Ambragaeana, Callogaeana, Gaeana, Sulphogaeana and Trengganua. Also similar to Balinta, Paratalainga, and Talainga.

Vittagaeana dives (Westwood, 1842), formerly Tosena dives:
Tosena dives (Westwood, 1842)
More info: Vittagaeana dives (Westwood, 1842)

Vittagaeana paviei (Noualhier, 1896), formerly Tosena paviei:
Tosena paviei
More info: Vittagaeana paviei (Noualhier, 1896)

December 28, 2020

Cicada Research Published in 2020

Filed under: Papers and Documents — Dan @ 11:46 pm

2020 was quite a year. In spite of everything going on in the world, plenty of cicada research was published. Around 40 in all.

Below you will find a list organized by month. If I missed any, please email me at Here are the pages for 2019, 2018, and 2017.


Title: Cytogenetic characterization of periodical cicadas (Hemiptera: Cicadidae: Magicicada)
Authors: Gayane Karagyan, Natalia Golub, Teiji Sota
Journal: Eur. J. Entomol. 117: 474-480, 2020

Title: Phylogeography of Ryukyu insular cicadas: Extensive vicariance by island isolation vs accidental dispersal by super typhoon
Authors: Soichi Osozawa, Kenichi Kanai, Haruo Fukuda, John Wakabayashi
Journal: bioRxiv
Notes: Not peer reviewed yet.


Title: A new genus and three new species of South African Cicadettini (Hemiptera: Cicadidae: Cicadettinae)
Authors: Allen F. Sanborn, Martin H. Villet.
Journal: Zootaxa, Vol 4885, No 4.

Title: The cicadas (Hemiptera: Cicadidae) of Ecuador including the description of five new species, a new subtribe, four new synonymies, and fifteen new records
Author: Allen F. Sanborn
Journal: Zootaxa Vol 4880, No 1

Title: The cicadas (Hemiptera: Cicadidae) of Suriname including the description of two new species, five new combinations, and three new records
Author: Allen F. Sanborn
Journal: Zootaxa Vol 4881, No 3


Title: A revision of the Australian cicada genus Punia Moulds, 2012 (Cicadidae: Cicadettinae: Cicadettini) with the description of four new species.
Author: M.S. Moulds.
Journal: Zootaxa, Vol 4860, No 1.

Title: A new species of Okanagana from the Walker Lane region of Nevada and California (Hemiptera: Auchenorrhyncha: Cicadidae)
Author: Will Chatfield-Taylor, Jeffrey A. Cole
Journal: Zootaxa, Vol 4868, No 4

Title: A New Species of Herrera Distant, 1905 (Hemiptera: Cicadoidea: Cicadidae: Cicadettinae: Carinetini) from Panama with New Records for Two Additional Species
Author: Allen F. Sanborn
Journal: Proceedings of the Entomological Society of Washington, 122(4):907-915 (2020).

Title: Checklist and provisional atlas of singing cicadas (Hemiptera: Cicadidae) of Bulgaria, based on bioacoustics
Author: Tomi Trilar, Ilia Gjonov, Matija Gogala
Journal: Biodiversity Data Journal, 8.e54424


Title: N-Acetyldopamine derivatives from Periostracum Cicadae and their regulatory activities on Th1 and Th17 cell differentiation
Authors: Punam Thapaa, Ye Gua, Yun-Seo Kil, Su Cheol Baek, Ki Hyun Kim, Ah-Reum Han, Eun Kyoung Seo, Hyukjae Choi, Jae-Hoon Chang, Joo-Won Nam
Journal: Bioorganic Chemistry, Volume 102

Title: Two new species of Neotropical cicadas (Hemiptera: Cicadidae: Cicadettinae) from southeastern Brazil
Author: Allen F. Sanborn
Journal: Journal of Insect Biodiversity, Vol 19, No 1

Title: Predator avoidance leads to separate emergence cycles in the protoperiodical Okanagana magnifica Davis, 1919 (Hemiptera: Cicadidae)
Author: Will Chatfield-Taylor
Journal: The Pan-Pacific Entomologist, 96(3):135-141 (2020)

Title: Cicadas of Portugal, those musicians which give joy during summer: towards a better knowledge of their Natural History and Conservation
Author: J. A. Quartau
Journal: Naturae Digital, Revista de Cultura Científica do Museu Virtual da Biodiversidade da Universidade de Évora, 10 Setembro de 2020.
Link: Cigarras de Portugal, esses músicos estivais que animam o verão com as suas melopeias: para um melhor conhecimento da sua história natural e conservação.


Title: Ecological indices of phytophagous Hemiptera and their natural enemies on Acacia auriculiformis (Fabales: Fabaceae) plants with or without dehydrated sewage sludge application in a degraded area
Authors: Luan Rocha Dourado, Germano Leão Demolin-Leite, Marcus Alvarenga Soares, Gustavo Leal Teixeira, Farley William Souza Silva, Regynaldo Arruda Sampaio, Jose Cola Zanuncio, Jesusa Crisostomo Legaspi
Journal: PLoS ONE 15(8): e0237261

Title: The cicadas (Hemiptera: Cicadidae) of Trinidad and Tobago including the description of three new species and seven new records, with new records for several additional Caribbean Islands
Author: Allen F. Sanborn
Journal: Zootaxa Vol 4838, No 4

Title: Reassessment of the cicada fossil Oncotympana lapidescens Zhang, 1989 with the description of a new genus (Hemiptera, Cicadoidea, Cicadidae)
Author: M.S. Moulds
Journal: Palaeoentomology, Vol 3, No 4

Title: Standardisation of bioacoustic terminology for insects
Author: Edward Baker, David Chesmore
Journal: Biodiversity Data Journal, 8:e54222

Title: Four new species of cicadas in the Yoyetta abdominalis (Distant) species group (Hemiptera: Cicadidae: Cicadettinae) from southeastern Australia
Author: Lindsay W. Popple, David L. Emery
Journal: Records of the Australian Museum, 72(4): 123—147




Title: Behavioral betrayal: How select fungal parasites enlist living insects to do their bidding
Authors: Brian Lovett, Angie Macias, Jason E. Stajich, John Cooley, Jørgen Eilenberg, Henrik H. de Fine Licht, Matt T. Kasson
Journal: PLoS Pathog 16(6): e1008598

Title: Look closely and listen carefully: unexpected cicada diversity in northern Sardinia, with the description of a new species (Cicadidae: Tibicina)
Author: Thomas Hertach
Journal: Zoological Journal of the Linnean Society, zlaa047

Title: Additions to the cicada (Insecta: Hemiptera: Cicadidae) fauna
of India: first report and range extension of four species with notes on their natural history from Meghalaya

Authors: Vivek Sarkar, Cuckoo Mahapatra, Pratyush P. Mohapatra & Manoj V. Nair
Journal: Journal of Threatened Taxa, Vol. 12 | No. 9 | Pages: 16021—16042
DOI: 10.11609/jot.5668.12.9.16021-16042


Title: The cicadas (Hemiptera: Cicadidae) of Peru including the description of twenty-four new species, three new synonymies, and thirty-seven new records
Author: Allen F. Sanborn
Journal: Zootaxa Vol 4785, No 1

Title: Molecular and Topographical Organization: Influence on Cicada Wing Wettability and Bactericidal Properties
Authors: Jessica Román-Kustas, Jacob B. Hoffman, Julian H. Reed, Andrew E. Gonsalves, Junho Oh, Longnan Li, Sungmin Hong, Kyoo D. Jo, Catherine E. Dana, Nenad Miljkovic, Donald M. Cropek, Marianne Alleyne
Journal: Advanced Material Interfaces, Volume7, Issue10

Title: True cicadas (Cicadidae) as prey for the birds of the Western Palearctic: a review
Author: Pere Pons
Journal: Avian Research volume 11, Article number: 14 (2020)

Title: Evolutionary relationships among Massospora spp. (Entomophthorales), obligate pathogens of cicadas
Authors: Angie M. Macias, David M. Geiser, Jason E. Stajich, Piotr Lukasik, Claudio Veloso, DeAnna C. Bublitz, Matthew C. Berger, Greg R. Boyce , Kathie Hodge & Matt T. Kasson
Journal: Mycologia, Volume 112, 2020


Title: New genus and species of Leptopsaltriini (Hemiptera: Cicadidae: Cicadinae) from China and Vietnam, with colour-changing behaviour reported for the first time in Cicadoidea
Authors: Cong Wei, Siyue Wang, Masami Hayashi, Miao He, Hong Thai Pham
Journal: Zootaxa Vol 4759, No 2.

Title: Replacement name for the cicada genus Pycnoides Sanborn, 2020
(Hemiptera: Cicadidae: Cicadinae: Platypleurini) and sixteen new combinations

Author: Allen F. Sanborn
Journal: Zootaxa Vol 4759, No 4

Title: Eight new species and three new records of Neotropical cicadas (Hemiptera: Cicadidae) from Venezuela
Author: Allen F. Sanborn
Journal: Journal of Insect Biodiversity, Vol 16, No 1


Title: Redefinition of the cicada tribe Hemidictyini Distant, 1905, status of the tribe Iruanini Boulard, 1993 rev. stat., and the establishment of Hovanini n. tribe and Sapantangini n. tribe (Hemiptera: Cicadidae)
Authors: Allen F. Sanborn, David C. Marshall, Maxwell S. Moulds, Stéphane Puissant, Chris Simon
Journal: Zootaxa, Vol 4747, No 1.

Title: On the morphology and evolution of cicadomorphan tymbal organs
Authors: Leonidas-Romanos Davranoglou, Beth Mortimer, Graham K.Taylor, Igor Malenovský
Journal: Arthropod Structure & Development, Volume 55

Title: A new species of the genus Tanna Distant (Hemiptera: Cicadidae: Cicadinae: Leptopsaltriini) from Vietnam, with notes on taxonomic changes in Tanna
Authors: Young June Lee, David Lyall Emery
Journal: Journal of Asia-Pacific Biodiversity, Volume 13, Issue 1

Title: Evaluation of Organically Acceptable Methods to Control Periodical Cicada (Hemiptera: Cicadidae) Oviposition Injury on Nonbearing Apple Trees
Author: Daniel L. Frank
Journal: J. of Entomological Science, 55(2):210-218 (2020)


Title: Automated acoustic detection of a cicadid pest in coffee plantations
Authors: João Paulo Lemos Escola, Rodrigo Capobianco Guido, Ivan Nunes da Silva, Alexandre Moraes Cardoso, Douglas Henrique Bottura Maccagnan, Artur Kenzo Dezotti
Journal: Computers and Electronics in Agriculture, Volume 169.

Title: Quality consistency evaluation on four origins of Cicadae Periostracum by ultra-performance liquid chromatography coupled with quadrupole/time-of-flight mass spectrometry analysis
Authors: Xin-Ci Cao, Xiao-Ya Zhang, Jin-Di Xu, Hong Shen, Shan-Shan Zhou, He Zhu, Ming Kong, Wei Zhang, Gui-Rong Zhou, Yi He, Qian Mao, Song-Lin Li
Journal: Journal of Pharmaceutical and Biomedical Analysis, Volume 179

Title: Afromelampsalta, a new genus, a new species, and five new combinations of African cicadettine cicadas (Hemiptera: Cicadidae: Cicadettinae)
Authors: Allen F. Sanborn, Martin H. Villet
Journal: Zootaxa, Vol 4731, No 4.

Title: Emergence of a novel interaction between brown bear and cicada due to anthropogenic habitat modification
Authors: Kanji Tomita, Tsutom Hiura
Journal: bioRxiv
Notes: Not peer reviewed yet.

Title: Dyticodopoea and Pygmaeodopoea, New Genera for the Central American Cicada Species Previously Assigned to Odopoea Stål, 1861 (Hemiptera: Cicadoidea: Cicadidae: Cicadinae: Zammarini)
Author: Allen F. Sanborn
Journal: Proceedings of the Entomological Society of Washington, 122(1):117-126 (2020).


Title: A new species of Oligoglena Horváth, 1912 (Hemiptera: Cicadidae) from Mediterranean Turkey
Author: Abbas Mol.
Journal: Zoology in the Middle East

Title: Not all cicadas increase thermal tolerance in response to a temperature gradient in metropolitan Seoul
Authors: Hoa Quynh Nguyen, Hortense Serret, Yoonhyuk Bae, Seongmin Ji, Soyeon Chae, Ye Inn Kim, Jeongjoo Ha & Yikweon Jang
Journal: Sci Rep 10, 1343 (2020)

Title: Redescription of the cicada genus Pycna Amyot & Audinet-Serville, 1843 (Hemiptera: Cicadidae: Cicadinae: Platypleurini) with the formation of two new genera, one new species, one revised species status and twenty-four new combinations
Author: Allen F. Sanborn
Journal: Zootaxa, Vol 4722, No 2

No month

Title: Rapid authentication of the Cicadidae Periostracum using a COI sequence-based SCAR markers
Authors: Pureum Noh, Wook Jin Kim, Sungyu Yang, Inkyu Park, Jun-Ho Song, Goya Choi, Byeong Cheol Moon
Journal: Integrative Medicine Research, Volume 9, Supplement 1


Magicicada septendecim & Magicicada septendecula genome datasets including genome, transcript and protein sequence, annotation and a data report
Site: National Library of Medicine
Link: Magicicada genome datasets

Cicada Research Wordcloud

September 21, 2019

Cicada Papers Published in 2019

Filed under: Papers and Documents — Dan @ 5:00 am

This is a running list of papers or documents published about cicadas in the year 2019. 35 so far (as of November).

If I missed an article, email me at


  1. An augmented wood-penetrating structure: Cicada ovipositors enhanced with metals and other inorganic elements. Authors: Matthew S. Lehnert, Kristen E. Reiter, Gregory A. Smith & Gene Kritsky . Link to Nature.


  1. A revision of the Yoyetta abdominalis (Distant) species group of cicadas (Hemiptera: Cicadidae: Cicadettinae), introducing eight new species. Authors: David Emery, Nathan Emery, Lindsay Popple. Link to Journals Australia Museum.
  2. Off-target capture data, endosymbiont genes and morphology reveal a relict lineage that is sister to all other singing cicadas. Authors: Chris Simon, Eric R L Gordon, M S Moulds, Jeffrey A Cole, Diler Haji, Alan R Lemmon, Emily Moriarty Lemmon, Michelle Kortyna, Katherine Nazario, Elizabeth J Wade, Russell C Meister, Geert Goemans, Stephen M Chiswell, Pablo Pessacq, Claudio Veloso, John P Mccutcheon, Piotr Lukasik. Link to Biological Journal of the Linnean Society.
  3. Noisy neighbours among the selfish herd: a critical song distance mediates mate recognition within cicada emergences (Hemiptera: Cicadidae). Authors: Will Chatfield-Taylor, Jeffrey A Cole. Link to Biological Journal of the Linnean Society.
  4. Genital coupling, morphology and evolution of male holding structures in Cicadinae (Hemiptera: Cicadidae). Authors: Tatiana Petersen Ruschel, Filipe Michels Bianchi, Luiz Alexandre Campos. Link to Biological Journal of the Linnean Society.
  5. Psychoactive plant- and mushroom-associated alkaloids from two behavior modifying cicada pathogens. Authors: Greg R. Boyce, Emile Gluck-Thaler, Jason C. Slot, Jason E. Stajich, William J. Davis, Tim Y.James, John R. Cooley, Daniel G. Panaccione, Jørgen Eilenberg, Henrik H. De Fine Licht, Angie M. Macias, Matthew C. Berger, Kristen L. Wickert, Cameron M. Stauder, Ellie J. Spahr, Matthew D. Maust, Amy M. Metheny, Chris Simon, Gene Kritsky, Kathie T. Hodge, Richard A.Humber, Terry Gullion, Dylan P.G. Short, Teiya Kijimoto, Dan Mozgai, Nidia Arguedas, Matt T. Kasson. Link to


  1. A note on the taxonomy and natural history of the Summer Clicker Lahugada dohertyi (Distant, 1891) (Insecta: Hemiptera: Cicadidae) along with its distribution in northern West Bengal, India. Authors: Vivek Sarkar. Like to the Journal of Threatened Taxa.
  2. Gold coated Cicada wings: Anti-reflective micro-environment for plasmonic enhancement of fluorescence from upconversion nanoparticles. Authors: Akash Gupta, Hao-Yu Cheng, Kung-Hsuan Lin, Chien Ting Wu, Pradip Kumar Roy, Sandip Ghosh, Surojit Chattopadhyay. Link to Science Direct.
  3. A new species and first record of the cicada genus Sinotympana Lee, 2009 (Hemiptera: Cicadidae: Dundubiini) from Vietnam. Authors: THAI-HONG PHAM, ALLEN F. SANBORN, HUYEN-THI NGUYEN, JEROME CONSTANT. Link to Zootaxa.


  1. A New Species of Calopsaltria Stål, 1861 (Hemiptera: Cicadoidea: Cicadidae: Cicadettinae: Parnisini) from South Africa. Author: Allen F. Sanborn. Link to paper.
  2. The cicadas (Hemiptera: Cicadidae) of Bolivia including the descriptions of fifteen new species, the resurrection of one genus and two species, seven new combinations, six new synonymies, and twenty-eight new records. Author: ALLEN F. SANBORN. Link to Zootaxa.


  1. Periodical cicada emergence resource pulse tracks forest expansion in a tallgrass prairie landscape. Authors: Matt R. Whiles, Bruce A. Snyder, Brent L. Brock, Sophia Bonjour, Mac A. Callaham Jr., Clinton K. Meyer, Alex Bell. Link to Ecosphere journal.


  1. The cicada genus Selymbria Stå1, 1861 (Hemiptera: Cicadidae: Tibicininae: Selymbrini): redescription including ten new species and a key to the genus. Author: Allen F Sanborn. Link to Zootaxa.
  2. Cicada Endosymbionts Have tRNAs That Are Correctly Processed Despite Having Genomes That Do Not Encode All of the tRNA Processing Machinery. Authors: James T. Van Leuven, Meng Mao, Denghui D. Xing, Gordon M. Bennett, John P. McCutcheon. Link to mBio
  3. An enigmatic fossil hairy cicada (Hemiptera, Tettigarctidae) from mid-Cretaceous Burmese amber. Authors: Hui Jiang, Jun Chen, Ed Jarzembowski, Bo Wang. Link to article
  4. Comparative sialotranscriptome analysis of the rare Chinese cicada Subpsaltria yangi, with identification of candidate genes related to host-plant adaptation. Authors: Yunxiang Liu, Mengmeng Qi, Christopher H. Dietrich, Zhiqiang He, Cong Wei. Link to article


  1. First record of the cicadas genus Orientopsaltria Kato, 1944 (Hemiptera: Cicadidae) from Vietnam, with description of one new species. Authors: Thai-Hong Pham, Thi Huyen Nguyen, Jerome Constant. Link to Zootaxa


  1. The applications of biomimetic cicada-wing structure on the organic light-emitting diodes. Authors: Chih-Kai Nien, Hsin Her Yu. Link to Science Direct.
  2. Oxygen/phosphorus co-doped porous carbon from cicada slough as high-performance electrode material for supercapacitors. Authors: Bingwei Chen, Wenzhuo Wu, Chunyang Li, Yanfang Wang, Yi Zhang, Lijun Fu, Yusong Zhu, Lixin Zhang & Yuping Wu. Link to Nature.
  3. The winner takes it all: how semelparous insects can become periodical. Authors: Odo Diekmann, Robert Planqué. Link to
  4. A new species of larval Caeculisoma (Acari: Erythraeidae) parasitic on cicadas from China with detailed comparison of all larval members in the genus. Authors: Si-Yuan Xu, Tian-Ci Yi, Jian-Jun Guo, and Dao-Chao Jin. Link to
  5. Characterization of polymorphic loci for two cicada species: Cryptotympana atrata and Hyalessa fuscata (Hemiptera: Cicadoidae). Author(s): Hoa Quynh Nguyen,Soyeon Chae, Erick Kim, Yikweon Jang. Link to paper.


  1. New Host Records and Biological Notes for Diceroprocta bulgara (Distant) in Mexico. Authors: José Antonio Sánchez-García , José Joaquín Velázquez-Monreal , Héctor Miguel Guzmán-Vásquez , Roselia Jarquín-López , Jesús Alberto Ortíz-López , Miguel Ángel Manzanilla-Ramírez , Manuel Ovando-Cruz , Teodulfo Aquino-Bolaños , Allen F. Sanborn. Link to paper.
  2. Cicada slough-derived heteroatom incorporated porous carbon for supercapacitor: Ultra-high gravimetric capacitance. Authors: Haiyang Jia, Jiawei Sun, Xiao Xie, Kuibo Yin, Litao Sun. Link to articles.
  3. Evolutionary hysteresis and ratchets in the evolution of periodical cicadas. Author(s): Jaakko Toivonen and Lutz Fromhage. Link to paper.
  4. The effects of pulsed fertilization and chronic herbivory by periodical cicadas on tree growth. Author(s): Louie H. Yang, Richard Karban. Link to paper.
  5. Mitochondrial Genomics Reveals Shared Phylogeographic Patterns and Demographic History among Three Periodical Cicada Species Groups. Author(s): Zhenyong Du, Hiroki Hasegawa, John R Cooley, Chris Simon, Jin Yoshimura, Wanzhi Cai, Teiji Sota, Hu Li. Link to paper.
  6. Homoptera — Cicadas and Hoppers. Author(s): Ying Wang, Xiao Zhang, Tingying Zhang, Xue Liu, Chungkun Shih, Yunzhi Yao, Dong Ren. Link to paper.
  7. Mesodiphthera Tillyard, 1919, from the Late Triassic of Queensland, the oldest cicada (Hemiptera: Cicadomorpha: Cicadoidea: Tettigarctidae). Author(s): KEVIN J. LAMBKIN. Link to paper.
  8. Out of Africa? A dated molecular phylogeny of the cicada tribe Platypleurini Schmidt (Hemiptera: Cicadidae), with a focus on African genera and the genus Platypleura Amyot & Audinet-Serville. Author(s): Benjamin W. Price, David C. Marshall, Nigel P. Barker, Chris Simon, Martin H. Villet. Link to paper.
  9. Ecophysiological responses to climate change in cicadas. Author(s): Minoru Moriyama , Hideharu Numata. Link to paper.


  1. Intra- and Interspecific Prey Theft in Cicada Killers (Hymenoptera: Apoidea: Sphecius) . Author(s): J R Coelho, C W Holliday, J M Hastings. Link to paper.


  1. Phylogeny and biogeography of the leaf-winged cicadas (Hemiptera: Auchenorrhyncha: Cicadidae). Author(s): Tatiana Petersen Ruschel, Luiz Alexandre Campos. Link to paper.
  2. A Simple Model of Periodic Reproduction: Selection of Prime Periods. Author(s): Raul Abreu de Assis, Mazílio Coronel Malavazi. Link to paper.
  3. First hairy cicadas in mid-Cretaceous amber from northern Myanmar (Hemiptera: Cicadoidea: Tettigarctidae). Author(s): Yanzhe Fu, Chenyang Cai, Diying Huang. Link to paper.
  4. Investigation of nanostructure-based bactericidal effect derived from a cicada wing by using QCM-D. Authors: Keisuke Jindai, Kazuki Nakadea, Takashi Sagawa, Hiroaki Kojima, Tomohiro Shimizu, Shoso Shingubara, Takeshi Ito. Link to Science Direct.
  5. Hemiptera of Canada. Authors: Robert G. Foottit, H. Eric L. Maw, Joel H. Kits, Geoffrey G. E. Scudder. Link to Zookeys.

March 23, 2019

Platypleura hampsoni (Distant, 1887)

Filed under: India | Oriental Cicadidae | Platypleura | Platypleurini | W. L. Distant — Dan @ 1:01 am

Platypleura hampsoni is a cicada found in India.

Platypleura hampsoni

Image and Description from A Monograph of Oriental Cicadas by W. L. Distant. 1889-1892. Read it on the Biodiversity Heritage Library website:

Male. Head luteous; front with a number of black linear markings; vertex with a transverse, narrow, black fascia between the eyes, and with a central black spot containing the ocelli. Pronotum greenish-ochraceous, the disk with the following black markings: — a central I-shaped spot, on each side of which are some oblique linear markings; the lateral dilated margins are black, and the anterior margin is narrow — and the posterior margin broadly— dull reddish ochraceous. Mesonotum greenish-ochraceous, with the following black spots: — four obconical from anterior margin, of which the central two are smallest; and a large, oblong, discal spot, with a small partly rounded spot on each side of it; the basal cruciform elevation dull reddish ochraceous. Abdomen above black. Head beneath, with the face black, marked with luteous transverse lines; sternum somewhat ochraceously pilose; abdomen beneath black, the segmental margins ochraceous, the anal appendage of the same color; legs castaneous, streaked or spotted with piceous and luteous. Rostrum black, the basal portion luteous.

Tegmina pale hyaline, with the venation brown, the costal membrane greenish, the basal third somewhat opaque, with darker transverse markings and small basal black markings; a double irregular series of dark brown spots cross the tegmina at about center, a dark brown fascia at bases of upper apical areas, a few small subapical spots and some small marginal spots of the same color. Wings brownish-ochraceous, paler at apex than at base and very pale across the center, with a white marginal spot near anal angle; the venation brown.

The rostrum reaches the basal abdominal segment; the lateral margins of the pronotum are distinctly angulated; the face is robustly gibbous, with a profound central longitudinal sulcation; the posterior tibijE have three distinct spines on each side of apical half.

Long. excl. tegm. 2 . 23 millim. Exp. tegm. 70 millim. ; exp. pronot. angl. 13 millim.

Hab. — Continental India : Neelgiri Hills, northern slopes, 3500 & 5000 feet (Hampson — coll. Dist.).

Scientific classification:
Family: Cicadidae
Subfamily: Cicadinae
Tribe: Platypleurini
Genus: Platypleura
Species: Platypleura hampsoni (Distant, 1887)

For more information about this cicada, visit Cicadas of India.

February 27, 2019

Tosena albata Distant, 1878

Filed under: Burma | India | Michel Chantraine | Nepal | Oriental Cicadidae | Thailand | Tosena | Tosenini | Vietnam | W. L. Distant — Tags: — Dan @ 1:01 am

Tosena albata is a cicada found in Burma, India, Nepal, Thailand, and Vietnam. In the past, this cicada was known as Tosena melanoptera.

Photo by Michel Chantraine:
Tosena albata


Tegmina with five longitudinal greyish fasciae, one occupying radial area, and the others divided by ulnar veins, two of which amalgamate with the transverse fascia, and five irregular greyish fasciae situate in the apical areas. Wings with nine greyish ray-like fasciae situate on outer margin, the one at anal area largest; two-thirds of the outer margin narrowly edged with greyish. Apices of the femora not ochraceous.

Long. excl. tegm. Male 58 to 62 millim.; Female 48 millim. Exp. tegm. Male, 140 to 155 millim. Female, 130 millim.

Scientific classification:
Family: Cicadidae
Subfamily: Cicadinae
Tribe: Tosenini
Genus: Tosena
Species: Tosena albata Distant, 1878

For more information about this cicada, visit Cicadas of India.


  1. The description and location information comes from A Monograph of Oriental Cicadas by W. L. Distant. 1889-1892. Read it on the Biodiversity Heritage Library website.
  2. Species name information comes from Allen Sanborn’s Catalogue of the Cicadoidea (Hemiptera: Auchenorrhyncha).

February 21, 2019

Distantalna splendida splendida (Distant, 1878)

Distantalna splendida splendida is a cicada found in China, Thailand, India, Burma, Cambodia, Laos, and Vietnam. It is formerly known as Tosena splendida. It is known in pop culture as the basis for the Cicada 3301 logo.

Photo by Dan Mozgai:
Distantalna splendida formerly Tosena splendida

Photo by Michel Chantraine:

This description comes from A Monograph of Oriental Cicadas by W. L. Distant. 1889-1892. Read it on the Biodiversity Heritage Library website:

Body above black; front with a reddish spot at each basal angle; eyes, two small spots on vertex, four large spots to pronotum (two on disk and one at each posterior lateral angle) and two spots on disk of mesonotum,* luteous. Body beneath and legs black; lateral margins of the face, a wide central annulation to femora, and a central discal series of subtriangular spots, sanguineous.

Tegmina and wings— where not obscured by darker markings— pale hyaline, exhibiting varied opaline luster, which in some lights is found to be ornamented with close and regular series of transverse darker strife ; tegmina at base (narrowly) and costal membrane shining blackish ; venation bright luteous and for two-thirds from base broadly margined with shining blackish, and a series of shining blackish marginal spots on the apices of longitudinal veins to apical areas largest and somewhat fused at apex; claval area pale greenish. Wings pale greenish for nearly two-thirds their area from the base; remaining apical area shining blackish, enclosing a submarginal series of pale opaline spots, of which the largest are subapical.

Scientific classification:
Family: Cicadidae
Subfamily: Cicadinae
Tribe: Ayuthiini (formerly Tosenini as of 20212)
Subtribe: Distantalnina
Genus: Distantalna
Species: Distantalna splendida splendida (Distant, 1878)


  1. Species name change information comes from Allen Sanborn’s Catalogue of the Cicadoidea (Hemiptera: Auchenorrhyncha).
  2. New Tribe: Hill Kathy B. R., Marshall David C., Marathe Kiran, Moulds Maxwell S., Lee Young June, Pham Thai-Hong, Mohagan Alma B., Sarkar Vivek, Price Benjamin W., Duffels J. P., Schouten Marieke A., de Boer Arnold J., Kunte Krushnamegh, Simon Chris (2021) The molecular systematics and diversification of a taxonomically unstable group of Asian cicada tribes related to Cicadini Latreille, 1802 (Hemiptera:Cicadidae). Invertebrate Systematics 35, 570-601.

February 18, 2019

Cryptotympana aquila (Walker, 1850)

Cryptotympana aquila is a cicada found in Indonesia, Malaysia, Brunei, Thailand, and likely adjacent nations.

Photo by Michel Chantraine:
Photo by Michel Chantraine

This description comes from A Monograph of Oriental Cicadas by W. L. Distant. 1889-1892. Read it on the Biodiversity Heritage Library website.

Male: Body above black; eyes ochraceous; ocelli luteous. Pronotum with some indistinct discal markings and the posterior margin pale castaneous. Mesonotum with a central triangular linear fascia, on each side of which is an irregular and inwardly notched fascia, and the basal cruciform elevation, pale castaneous. Abdomen black, the lateral margins fringed with pale castaneous pile. Body beneath very pale castaneous; the face, anterior margin of the head between face and eyes, and basal abdominal patch, black. Anterior legs black, the femera streaked with pale castaneous; intermediate legs with the femora black streaked with pale castaneous, the tibis pale castaneous with their bases and apices black; posterior legs pale castaneous, the bases and apices of femora and tibiae black.

Tegmina and wing pale hyaline; tegmina with the venation and the costal membrane pale castaneous, the extreme basal margin of the last black; the basal third (excluding venation) and the subcostal area to apex, blackish; wings with about basal half obliquely black.

The body is robust but moderately elongate; the opercula do not overlap at their basal margins, and at a short distance from base become widely divergent and narrowed to apices, their outer margins slightly concavely sinuate and in length, they extend a little beyond the middle of the abdomen.

Long. excl. tegm. Male, 40 to 45 millim. Exp. tegm. 115 to 120 millim.

Scientific classification:
Family: Cicadidae
Subfamily: Cicadinae
Tribe: Cryptotympanini
SubTribe: Cryptotympanina
Genus: Cryptotympana
Species: Cryptotympana aquila (Walker, 1850)

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