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Cicada Mania France Lyristes Tacuini (Cryptotympanini) Tibicen

French Cicada

Cicada Mania has been around since 1996. It’s lived on at least five different domains (cicadamania.com is the current domain). Lots of history. Yesterday I was looking at old versions of the site on archive.org’s Wayback machine, which created archives of websites. I came across this post from 2001 and thought “these photos are not on the current site, let me upload them.

So here’s a copy of the original post, approximately 18 years later.

BTW, the cicada is Lyristes/Tibicen plebejus.

French Cicada

Cicada Experts, try to ID this French cicada!

Hello, In the south of France last week I saw a big insect. From the moment it climbed up a flower (picture 1) till the moment it became the one of picture 2 (10 hours later) I made 300 pictures with my Sony Mavica. Some questions; a) is this a cicada b) if so, whats the name c) do you know a site where I can find the sound of this species d) who could be interested in my pictures of the complete metamorphosis Greetings, H. Bakkenes Holland.

Tibicen/Lyristes plebejus

Tibicen/Lyristes plebejus

[Adding the old “back to top” cicada for fun.]

Back to top

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Lyristes Megatibicen Neotibicen Tibicen Young June Lee

Gigatibicen, Ameritibicen, Paratibicen

Gigatibicen
For some reason I associate “Giga” with “Gigabytes” and storage media like Flash drives, which explains this joke image.

Over the past two years there have been quite a few updates to the genera of the cicadas that were organized under the Tibicen genus earlier this decade.

The most recent paper by Young June Lee introduces the genera Gigatibicen, Ameritibicen, and Paratibicen 1. Earlier this year there was a paper by Allen Sanborn and Maxine Heath that introduced the genus Megatibicen 3, and in 2015 there was a paper by Kathy Hill and others that introduced Neotibicen and Hadoa 2.

See the end of the article for links to these papers, and related articles on CicadaMania.com.

The table below shows the names/synonyms (sub species have been removed to keep the table compact):

(Tibicen circa 2014) Hill (2015)2 Sanborn, Heath (2016)3 Lee (2016)1
Tibicen auletes Neotibicen auletes Megatibicen auletes Gigatibicen auletes
Tibicen auriferus Neotibicen auriferus no change no change
Tibicen canicularis Neotibicen canicularis no change no change
Tibicen cultriformis Neotibicen cultriformis Megatibicen cultriformis Ameritibicen cultriformis
Tibicen davisi davisi Neotibicen davisi no change no change
Tibicen dealbatus Neotibicen dealbatus Megatibicen dealbatus Ameritibicen dealbatus
Tibicen dorsatus Neotibicen dorsatus Megatibicen dorsatus Ameritibicen dorsatus
Tibicen figuratus Neotibicen figuratus Megatibicen figuratus Ameritibicen figuratus
Tibicen latifasciatus Neotibicen latifasciatus no change no change
Tibicen linnei Neotibicen linnei no change no change
Tibicen lyricen engelhardti Neotibicen lyricen no change no change
Tibicen pronotalis pronotalis Neotibicen pronotalis Megatibicen pronotalis Ameritibicen pronotalis
Tibicen pruinosus fulvus Neotibicen pruinosus no change no change
Tibicen resh Neotibicen resh Megatibicen resh Gigatibicen resh
Tibicen resonans Neotibicen resonans Megatibicen resonans Gigatibicen resonans
Tibicen robinsonianus Neotibicen robinsonianus no change no change
Tibicen similaris Neotibicen similaris no change Paratibicen similaris
Tibicen superbus Neotibicen superbus no change no change
Tibicen tibicen australis Neotibicen tibicen no change no change
Tibicen tremulus Neotibicen tremulus Megatibicen tremulus Ameritibicen tremulus
Tibicen winnemanna Neotibicen winnemanna no change no change
Tibicen bifidus Hadoa bifida no change no change
Tibicen chiricahua Hadoa chiricahua no change no change
Tibicen duryi Hadoa duryi no change no change
Tibicen inauditus Hadoa inaudita no change no change
Tibicen longioperculus Hadoa longiopercula no change no change
Tibicen neomexicensis Hadoa neomexicensis no change no change
Tibicen parallelus Hadoa parallela no change no change
Tibicen simplex Hadoa simplex no change no change
Tibicen texanus Hadoa texana no change no change
Tibicen townsendii Hadoa townsendii no change no change
  • 1 Lee, Y.J. 2016. Description of three new genera, Paratibicen, Gigatibicen, and Ameritibicen, of Cryptotympanini (Hemiptera: Cicadidae) and a key to their species Journal of Asia-Pacific Biodiversity, Volume 9, Issue 4, 1 December 2016, Pages 448—454. Link to Paper.
  • 2 Hill, et al. 2015. Molecular phylogenetics, diversification, and systematics of Tibicen Latreille 1825 and allied cicadas of the tribe Cryptotympanini, with three new genera and emphasis on species from the USA and Canada (Hemiptera: Auchenorrhyncha: Cicadidae), Zootaxa, Volume 3985, Issue 2, Pages 219—251. Link to Paper
  • 3 Sanborn A.F., Heath, M.S. 2016. Megatibicen n. gen., a new North American cicada genus (Hemiptera: Cicadidae: Cicadinae: Cryptotympanini), Zootaxa, Volume 4168, Issue 3. Link to Paper
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Auritibicen Chris Simon David Marshall Hadoa Kathy Hill Lyristes Max Moulds Neotibicen Tibicen

Major Changes to the Tibicen genera

Sometimes you wake up and the whole world is different. See this cicada:

Teneral Neotibicen tibicen
photo by me.

… when I went to sleep she was a Tibicen tibicen tibicen, but now I know she is a Neotibicen tibicen tibicen. 10 years ago, she was a Tibicen chloromera. 130 years ago, she was Cicada tibicen. Cicada names change as researchers discover their differences.

Two new papers have split the Tibicen (or Lyristes) genera into many genera: Tibicen (European Tibicen), Auritibicen (Tibicen of Asia/Japan), Neotibicen (mostly eastern North American Tibicen), and Hadoa (Tibicen of the western United States).

The first paper is Description of a new genus, Auritibicen gen. nov., of Cryptotympanini (Hemiptera: Cicadidae) with redescriptions of Auritibicen pekinensis (Haupt, 1924) comb. nov. and Auritibicen slocumi (Chen, 1943) comb. nov. from China and a key to the species of Auritibicen by Young June Lee, 2015, Zootaxa 3980 (2): 241—254. This paper establishes the new genera Auritibicen, and the members of the Tibicen/Lyristes genera fall into that genera. Here is a link. So, Tibicen flammatus aka Lyristes flammatus of Japan, for example, becomes Auritibicen flammatus.

A. flammatus
Auritibicen flammatus photo by Osamu Hikino.

The second paper is Molecular phylogenetics, diversification, and systematics of Tibicen Latreille 1825 and allied cicadas of the tribe Cryptotympanini, with three new genera and emphasis on species from the USA and Canada (Hemiptera: Auchenorrhyncha: Cicadidae) by Kathy B. R. Hill, David C. Marshall, Maxwell S. Moulds & Chris Simon. 2015, Zootaxa 3985 (2): 219—251. This paper establishes the Neotibicen (Hill and Moulds), and Hadoa (Moulds) genera. This paper also sought to establish the Subsolanus genera for the Asian Tibicen/Lyristes species but the previously mentioned paper by Young June Lee has precedence because it was published first. Link to paper.

To recap, European Tibicen/Lyristes are Tibicen

Lyristes plebejus photo by Iván Jesús Torresano García
Tibicen plebejus photo by Iván Jesus Torresano García.

… Asian Tibicen/Lyristes are now Auritibicen. Mostly-eastern North American Tibicen are now Neotibicen, and Western North American Tibicen are now Hadoa. Note that, the catagorization is not due to location, but to genetic and physiological evaluation (read the papers).

Needless to say this website and others have a lot of name changing to do, but in the mean time, here’s where the North American species fall out:

Neotibicen
Tibicen auletes Neotibicen auletes
Tibicen auriferus Neotibicen auriferus
Tibicen canicularis Neotibicen canicularis
Tibicen cultriformis Neotibicen cultriformis
Tibicen davisi davisi Neotibicen davisi davisi
Tibicen davisi harnedi Neotibicen davisi harnedi
Tibicen dealbatus Neotibicen dealbatus
Tibicen dorsatus Neotibicen dorsatus
Tibicen figuratus Neotibicen figuratus
Tibicen latifasciatus Neotibicen latifasciatus
Tibicen linnei Neotibicen linnei
Tibicen lyricen engelhardti Neotibicen lyricen engelhardti
Tibicen lyricen lyricen Neotibicen lyricen lyricen
Tibicen lyricen virescens Neotibicen lyricen virescens
Tibicen pronotalis pronotalis Neotibicen pronotalis pronotalis
Tibicen pronotalis walkeri Neotibicen pronotalis walkeri
Tibicen pruinosus fulvus Neotibicen pruinosus fulvus
Tibicen pruinosus pruinosus Neotibicen pruinosus pruinosus
Tibicen resh Neotibicen resh
Tibicen resonans Neotibicen resonans
Tibicen robinsonianus Neotibicen robinsonianus
Tibicen similaris Neotibicen similaris
Tibicen superbus Neotibicen superbus
Tibicen tibicen australis Neotibicen tibicen australis
Tibicen tibicen tibicen Neotibicen tibicen tibicen
Tibicen tremulus Neotibicen tremulus
Tibicen winnemanna Neotibicen winnemanna
Hadoa
Tibicen bifidus Hadoa bifida
Tibicen chiricahua Hadoa chiricahua
Tibicen duryi Hadoa duryi
Tibicen inauditus Hadoa inaudita
Tibicen longioperculus Hadoa longiopercula
Tibicen neomexicensis Hadoa neomexicensis
Tibicen parallelus Hadoa parallela
Tibicen simplex Hadoa simplex
Tibicen texanus Hadoa texana
Tibicen townsendii Hadoa townsendii
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Lyristes Tibicen

Tibicen or Lyristes

A Tibicen by any other name would still sound as sweet…

I always wondered why Lyristes plebejus is also called Tibicen plebejus.

It seems that there is a dispute as to whether the genus Tibicen should actually be called Lyristes. A petition was made (back in the 1980s) to the International Commission on Zoological Nomenclature, to change Tibicen to Lyristes. I learned this from the wonderful new book, The Cicadas (Hemiptera: Cicadoidea: Cicadidae) of North America North of Mexico by Allen F. Sanborn and Maxine S. Heath (order it). I checked the ICZN website, and the petition appears to fallen off their docket of open cases. I also noticed that on European and Japanese websites, they use Lyristes.

I personally hope the genus name doesn’t change for North American species — I would have to make a lot of changes on this website. Going through the name change from Tibicen chloromera to Tibicen chloromerus to Tibicen tibcen, was bad enough.

The root of the word Tibicen is flute player, and the root of the word Lyristes is lyre — both referring to musical instruments. (Frankly I think most Tibicen sound like power tools — I don’t know Latin for Black & Decker).

BTW, this is a Lyristes plebejus (from Spain):
Lyristes plebejus photo by Iván Jesús Torresano García
Photo by Iván Jesús Torresano García.

And this is a Lyristes flammatus (from Japan):
A. flammatus
Photo by Osamu Hikino

And some day, this might be a Lyristes auletes (from North Carolina):
Neotibicen auletes found in Winston-Salem, NC by Erin Dickinson. 2011.
Photo by Erin.

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Auritibicen Graptopsaltria Japan Lyristes Osamu Hikino Platypleura Tanna

Cicadas from Japan

I re-scanned some old (10+ years old) photos from Osamu Hikino.

Graptopsaltria nigrofuscata:

Graptopsaltria nigrofuscata

Platypleura kaempferi (Fabricius, 1794):

Platypleura kaempferi (Fabricius, 1794)

Amazing camouflage!

A male Tanna japonensis:

A male Tanna japonensis

A male Auritibicen japonicus:

Male Auritibicen japonicus (formerly Tibicen japonicus, Lyristes japonicus)

A male Auritibicen japonicus:

Male Auritibicen japonicus (formerly Tibicen japonicus, Lyristes japonicus)

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Ivan Garcia Lyristes Spain Video

More cicadas from Spain

Iván Jesus Torresano García sent us another 44 photos of Tibicen plebejus aka Lyristes plebejus, and several YouTube videos.

31 Tibicen plebejus molt sequence, photos by Iván Jesus Torresano García

Some YouTube videos:

Momento en el que finaliza de emerger Cicada 27/June/2007.

Tibicen plebejus recién cogida del parque Forestal Entrevías:

Tibicen plebejus recogida del parque.Emergiendo.27/June/2007

Lyristes plebejus or Tibicen plebejus?Emerging 27/June/2007.

Here’s a cicada news cast from Spain:

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Ivan Garcia Lyristes Spain

Cicada from Spain

We have a new gallery, our first from Spain. Tibicen plebejus by Iván Jesus Torresano García. 51 photos in all.

Here’s a sample:

Cicada from Spain