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November 24, 2021

Cicada Research Published in 2021

Filed under: Papers and Documents — Dan @ 10:36 pm

This is a list of cicada-related research published in 2021.

More than 60 so far (12/24/2021). Email with any more.


  1. Dandan, Wang & Huang, Zhi & Billen, Johan & Zhang, Guoyun & He, Hong & Wei, Cong. (2021). Complex co-evolutionary relationships between cicadas and their symbionts. Environmental Microbiology. 10.1111/1462-2920.15829. Link.
  2. Kritsky, Gene. (2021). One for the Books: The 2021 Emergence of the Periodical Cicada Brood X. American Entomologist. 67. 40-46. 10.1093/ae/tmab059. Link.
  3. Tomita, Kanji & Makoto, Kobayashi. (2021). Development of experimental mesocosms for cicada nymphs Graptopsaltria nigrofuscata: methodology and research recommendations. 93. 207-212. 10.25674/so93iss3id171. Link.
  4. Moulds, Max. (2021). A revision of the endemic Australian cicada genus Jassopsaltria Ashton, 1914 (Cicadidae: Cicadinae: Jassopsaltriini). Australian Entomologist. 48. 217-240. Link.
  5. Moriyama, Minoru & Yasuyama, Kouji & Numata, Hideharu. (2021). The formation of a hatching line in the serosal cuticle confers multifaceted adaptive functions on the eggshell of a cicada. Zoological Letters. 7. 10.1186/s40851-021-00178-8. Link.
  6. Tomita, Kanji. (2021). Camera traps reveal interspecific differences in the diel and seasonal patterns of cicada nymph predation. The Science of Nature. 108. 10.1007/s00114-021-01762-w. Link.
  7. Tian, Lejie & Liu, Hui & Zhang, Beiyi & Liu, Yi & Lv, Siye & Pang, Lingyan & Li, Junqi. (2021). Ni and CeO 2 Nanoparticles Anchored on Cicada-Wing-like Nitrogen-Doped Porous Carbon as Bifunctional Catalysts for Water Splitting. ACS Applied Nano Materials. 10.1021/acsanm.1c03850. Link.
  8. Sanborn, Allen & Cole, Jeffrey & Stukel, Mark & Lukasik, Piotr & Veloso, Claudio & Gonzalez, Valorie & Karkar, Jessica & Simon, Chris. (2021). Thirteen new species of Chilecicada Sanborn, 2014 (Hemiptera: Auchenorrhyncha: Cicadidae: Tibicininae) expand the highly endemic cicada fauna of Chile. Zootaxa. 5078. 1-70. 10.11646/zootaxa.5078.1.1. Link.


  1. Kojevnikova, A.. (2021). The Study of Cicadas in the Conditions of the Fergana Valley of Uzbekistan. International Journal of Multidisciplinary Research and Analysis. 04. 10.47191/ijmra/v4-i11-38. Link.
  2. Huang, Ailin & Wu, Tao & Wu, Xiuyun & Zhang, Biao & Shen, Yuanyuan & Wang, Suying & Song, Wenjun & Ruan, Haihua. (2021). Analysis of Internal and External Microorganism Community of Wild Cicada Flowers and Identification of the Predominant Cordyceps cicadae Fungus. Frontiers in microbiology. 12. 752791. 10.3389/fmicb.2021.752791. Link.
  3. Mozaffarian, Fariba. (2021). Cicadas and hoppers as pests of field crops in Iran (Hemiptera: Auchenorrhyncha). Link.
  4. Mukaimine, Wataru & Kawatsu, Kazutaka & Toquenaga, Yukihiko. (2021). Digging out intersexual and meteorological effects on cicada emergence using 10-year citizen monitoring. Ecological Entomology. 10.1111/een.13109. Link.
  5. Poerio, Aurelia & Girardet, Thomas & Petit, Chloé & Fleutot, Solenne & Jehl, Jean-Philippe & Arab-Tehrany, Elmira & Mano, João F. & Cleymand, Franck. (2021). Comparison of the Physicochemical Properties of Chitin Extracted from Cicada orni Sloughs Harvested in Three Different Years and Characterization of the Resulting Chitosan. Applied Sciences. 11. 11278. 10.3390/app112311278. Link.


  1. Lee, Young & Mohagan, Alma. (2021). A new species of Oncotympana Stål, 1870 (Hemiptera: Cicadidae: Oncotympanini) from Negros, Philippines. Journal of Asia-Pacific Biodiversity. Link.
  2. Simon, Chris & Cooley, John & Karban, Richard & Sota, Teiji. (2021). Advances in the Evolution and Ecology of 13- and 17-Year Periodical Cicadas. Annual review of entomology. 67. 10.1146/annurev-ento-072121-061108. Link.
  3. Nagata, Nobuaki & Toda, Mamoru & Ohbayashi, Takashi & Hayashi, Masami & Sota, Teiji. (2021). Phylogeography of cicadas on continental and oceanic islands in the northwestern Pacific region. Journal of Biogeography. 48. 10.1111/jbi.14262. Link.
  4. Umam, Rofiqul & Kinarya P, Endah & Yakin, Khusnul & Iqbal, Mochamad & Siregar, Rahmat nawi & Husein, Irzaman. (2021). Intensity level as sustainable energy: Analysis of the conversion of energy stored on cicadas sound waves. Jurnal Ilmiah Pendidikan Fisika Al-Biruni. 10. 35-44. 10.24042/jipfalbiruni.v10i2.9798. Link.


  1. Moulds, Max & Marshall, D & Popple, Lindsay. (2021). Kimberpsaltriini, a new tribe for a new Australian cicada allied to Talcopsaltria Moulds (Hemiptera: Cicadoidea: Cicadidae). Australian Entomologist. 48. 149-160. Link.
  2. Zhang, Lijia & Wang, Siyue & Billen, Johan & Wei, Cong. (2021). Morphology and ultrastructure of the epithelial femoral gland in cicadas (Hemiptera: Cicadidae). Arthropod Structure & Development. 64. 101086. 10.1016/j.asd.2021.101086. Link.
  3. Hajong, Sudhanya & Limatemjen,. (2021). Platylomia kohimaensis n. sp.—a new cicada species (Hemiptera: Cicadidae) from the Naga Hills in the Eastern Himalayas. Zootaxa. 5047. 081-091. 10.11646/zootaxa.5047.1.7. Link.
  4. Kalesh, S.. (2021). A new species of Pomponia Stål, 1866 (Hemiptera: Cicadidae) from the Western Ghats, with notes on the status of P. linearis (Walker, 1850) from southern India. Zootaxa. 5040. 388-403. 10.11646/zootaxa.5040.3.4. Link.
  5. Bator, John & Marshall, David & Hill, Kathy & Cooley, John & Leston, Adam & Simon, Chris. (2021). Phylogeography of the endemic red-tailed cicadas of New Zealand (Hemiptera: Cicadidae: Rhodopsalta ), and molecular, morphological and bioacoustical confirmation of the existence of Hudson’s Rhodopsalta microdora. Zoological Journal of the Linnean Society. 10.1093/zoolinnean/zlab065. Link.
  6. Zhang, Lijia & Wang, Siyue & Billen, Johan & Wei, Cong. (2021). Morphology and ultrastructure of the epithelial femoral gland in cicadas (Hemiptera: Cicadidae). Arthropod Structure & Development. 64. 101086. 10.1016/j.asd.2021.101086. Link.
  7. Nguyen, Hoa & Kim, Yuseob & Yikweon, Jang. (2021). De Novo Transcriptome Analysis Reveals Potential Thermal Adaptation Mechanisms in the Cicada Hyalessa fuscata. Animals. 11. 2785. 10.3390/ani11102785. Link.
  8. Qian, Yi & Sun, Xin & Wang, Xin & Yang, Xin & Fan, Mengyao & Zhong, Jiao & Pei, Zejun & Guo, Junping. (2021). Mechanism of Cordyceps Cicadae in Treating Diabetic Nephropathy Based on Network Pharmacology and Molecular Docking Analysis. Journal of Diabetes Research. 2021. 1-10. 10.1155/2021/5477941. Link.


  1. Villet, Martin & Edwards, Shelley. (2021). The cicada genus Tugelana Distant, 1912 (Hemiptera, Cicadidae): phylogenetic position and conservation status. African Invertebrates. 62. 399-410. 10.3897/afrinvertebr.62.66891. Link.
  2. Haji, Diler & Vailionis, Jason & Stukel, Mark & Gordon, Eric & Lemmon, Emily & Lemmon, Alan & McCutcheon, John & Simon, Chris. (2021). Host-associated microbial diversity in New Zealand cicadas uncovers elevational structure and replacement of obligate bacterial endosymbionts by Ophiocordyceps fungal pathogens. 10.1101/2021.08.24.457591. Link.


  1. Zhao, Chenchen. (2021). High-throughput sequencing yields a complete mitochondrial genome of the Cryptotympana atrata (Insecta: Hemiptera: Cicadidae). Mitochondrial DNA Part B. 6. 1883-1885. 10.1080/23802359.2021.1934154. Link.
  2. Pham, Thai & Lee, Young June. (2021). A new genus and species of the subtribe Leptopsaltriina (Hemiptera: Cicadidae: Leptopsaltriini) from Vietnam, with a key to the genera of Leptopsaltriina. Journal of Asia-Pacific Entomology. 10.1016/j.aspen.2021.07.011. Link.
  4. Sousa P, Grosso-Silva JM, Andrade R, Chaves C, Pinto J, Paupério J, Beja P, Ferreira S (2021) The InBIO Barcoding Initiative Database: DNA barcodes of Portuguese Hemiptera 01. Biodiversity Data Journal 9: e65314. Link.
  5. Malakoff, David & Stokstad, Erik. (2021). Something is killing U.S. birds. It’s not cicadas. Science. 373. 146-146. 10.1126/science.373.6551.146. Link.
  6. Kojevnikova, Alevtina. (2021). HARMFUL CICADA GENUS EMPOASCA WALSH OF THE FERGANA VALLEY. Universum:Chemistry & biology. 84. 10.32743/UniChem.2021.84.6.11700. Link.
  7. Lao, Thuan & Trinh, Van Hanh & Vuong, Loi & Vu Tien, Luyen & Le, Thuy & Dinh, Hiep & Truong, Nguyen. (2021). Molecular record for the first authentication of Isaria cicadae from Vietnam. Open Life Sciences. 16. 711-718. 10.1515/biol-2021-0074. Link.


  1. Escola, João & Guido, Rodrigo & Silva, Ivan & Maccagnan, Douglas & Cardoso, Alexandre & Souza, Uender. (2021). Estado da arte no monitoramento acústico de Cicadidae em lavouras de café: State of the art in acoustic monitoring of Cicadidae in coffee crops. Revista Macambira. 5. e051007. 10.35642/rm.v5i1.562. Link.
  2. Emery, Nathan & Emery, David. (2021). A new cicada species, Haemopsalta eximia (Hemiptera: Cicadidae) from north-western Sydney, Australia. Australian Entomologist. 48. 97. Link.
  3. Sanborn, Allen & Allick, Earl & Apang, Sandee & Castillo, Izyanna & Cruz, Erica & Davis, Theophilus & Duncan, Cory & Fierro, Fanny & Gebaide, Marla & Luke, Abigail & Pacheco, Maria & Paz-Castillo, Daniel & Perez, Laura & Poeck, Ana & Seepersaud, Adrian & Valdes, Carolina. (2021). Flight system morphology and minimum flight temperature in North American cicadas (Insecta: Hemiptera: Cicadidae). Journal of Thermal Biology. 100. 103004. 10.1016/j.jtherbio.2021.103004. Link.
  4. Pons, Pere & Carbonell-Font, Rafael & Franch, Martí & Lay, Josep & Espejo Fraga, Daniel & Fontelles, Ferran & Funosas, David & Fusellas, Marc & Puig-Gironès, Roger & Tobella, Carles & Franch, Marc. (2021). Diversitat, distribució i fenologia de les cigales (Hemiptera: Cicadidae) a Catalunya (NE Península Iberica) – Diversity, distribution and phenology of cicadas (Hemiptera: Cicadidae) in Catalonia (NE Iberian Peninsula) – Butlletí de la Institució Catalana d’Història Natural. 85. 59. 10.2436/20.1502.01.74. Link.
  5. WANG, SIYUE & PENG, XIAODONG & Wei, Cong. (2021). A review of the cicada genus Haphsa Distant, 1905 with the description of one new species from China (Hemiptera: Cicadidae) and a phylogenetic analysis of Haphsa and its allies. Zootaxa. 4991. 523-538. 10.11646/zootaxa.4991.3.5. Link.
  6. Li J, Liu H, Wu Y, Ye L, Huang X (2021) A dataset on type specimens of hemipteran insects in China. Biodiversity Data Journal 9: e64443. Link.
  7. Crespi, Sarah. (2021). Cicada citizen science, and expanding the genetic code. Science. 10.1126/science.abj7361. Link.
  8. Tian, Juanjuan & Zhang, Cangping & Wang, XiaoMeng & Rui, Xin & Zhang, Qiuqin & Chen, Xiaohong & Dong, Mingsheng & Li, Wei. (2021). Structural characterization and immunomodulatory activity of intracellular polysaccharide from the mycelium of Paecilomyces cicadae TJJ1213. Food Research International. 147. 110515. 10.1016/j.foodres.2021.110515. Link.


  1. Dandan, Wang & Liu, Yunxiang & Yan, Su & Wei, Cong. (2021). Bacterial Communities in Bacteriomes, Ovaries and Testes of three Geographical Populations of a Sap-Feeding Insect, Platypleura kaempferi (Hemiptera: Cicadidae). Current Microbiology. 78. 10.1007/s00284-021-02435-7. Link.
  2. Cen, Qi-Wen & Wang, Zheng-Yun & Tang, Zhen-Xing & Zhang, Yu & Chen, Tao & Xue, Da-Wei & Xu, Ming-Feng & Bai, Xue-Lian & Zhou, Ting & Shi, Lu-E. (2021). Initial Purification of Antimicrobial Fermentation Metabolites from Paecilomyces cicadae and Its Antimicrobial Mechanism. LWT. 148. 111785. 10.1016/j.lwt.2021.111785. Link.
  3. Su, Qihui & Zhang, Zhicai & Liu, Xiaocui & Wang, Feng. (2021). The transcriptome analysis on urea response mechanism in the process of ergosterol synthesis by Cordyceps cicadae. Scientific Reports. 11. 10927. 10.1038/s41598-021-90377-2. Link.
  4. Zeng, Zhaoying & Mou, Dan & Luo, Li & Zhong, Wenlin & Duan, Lin & Xiao, Zou. (2021). Different Cultivation Environments Affect the Yield, Bacterial Community and Metabolites of Cordyceps cicadae. Frontiers in Microbiology. 12. 669785. 10.3389/fmicb.2021.669785. Link.


  1. EWART, A. & Moulds, Max. (2021). An impressive new species of Mugadina from Queensland with notes on a congeneric species (Cicadidae: Cicadettinae: Cicadettini). Zootaxa. 4963. 563-576. 10.11646/zootaxa.4963.3.9. Link.
  2. Cooley, John & Marshall, David & Simon, Chris. (2021). Documenting Single-Generation Range Shifts of Periodical Cicada Brood VI (Hemiptera: Cicadidae: Magicicada spp.). Annals of the Entomological Society of America. 114. 10.1093/aesa/saab007. Link.
  3. Sanborn, Allen. (2021). A new species, genus and tribe of cicada (Hemiptera: Cicadoidea: Cicadidae: Tibicininae) from Chile with a list of Chilean cicada fauna. Zootaxa. 4952. 87-100. 10.11646/zootaxa.4952.1.5. Link.
  4. Reid, Graham & McCormack, James & Habimana, Olivier & Bayer, Fabian & Goromonzi, Catherine & Casey, Eoin & Cowley, Aidan & Kelleher, Susan. (2021). Biomimetic Polymer Surfaces by High Resolution Molding of the Wings of Different Cicadas. Materials. 14. 1910. 10.3390/ma14081910. Link.


  1. WANG, JIALI & DUFFELS, JOANNES & WEI, CONG. (2021). Description of a new species of the genus Maua Distant (Hemiptera, Cicadidae) from China. Zootaxa. 4949. 582-588. 10.11646/zootaxa.4949.3.8. Link.
  2. Lee, Young June. (2021). A new species of Platypleura Amyot & Audinet-Serville (Insecta: Hemiptera: Cicadidae: Platypleurini) from Mindanao, Philippines. Journal of Asia-Pacific Biodiversity. 14. 10.1016/j.japb.2021.03.003. Link.
  3. Cole, Jeffrey. (2021). Bioacoustics of California Cacama Distant, 1904 with the first state record of C. moorei Sanborn & M. Heath, 2011 (Hemiptera: Auchenorrhyncha: Cicadidae). The Pan-Pacific Entomologist. 97. 10.3956/2021-97.1.13. Link.
  4. Pham, Thai & Lee, Young June. (2021). A new genus and species of the subtribe Aolina (Hemiptera: Cicadidae: Dundubiini) from Vietnam, with discussion on the taxonomic positions of Meimuna-like genera and species groups. Journal of Asia-Pacific Entomology. 24. 10.1016/j.aspen.2021.02.012. Link.


  1. Ruschel, Tatiana & Sanborn, Allen. (2021). Two new genera and species of Fidicinini Distant, 1905 with a re-description of Nosola Stål, 1866 (Hemiptera, Auchenorrhyncha, Cicadidae). Zootaxa. 4920. 509-527. 10.11646/zootaxa.4920.4.3. Link.
  2. Sanborn, Allen. (2021). The cicadas (Hemiptera: Cicadoidea: Cicadidae) of Madagascar including a new tribe, five new genera, twelve new species, four new species synonymies, five revised species status, ten new combinations, new tribal assignments for four genera, one new subtribe synonymy, a checklist and key to the species Zootaxa. Vol. 4937 No. 1: 24. Link.


  1. Thapa, Punam & Katila, Nikita & Choi, Hyukjae & Han, Ah-Reum & Choi, Dong-Young & Nam, Joowon. (2021). Neuroprotective Effects of N-Acetyldopamine Dimers from Cicadidae Periostracum. Link.
  2. Mahmoudikordi, Farzaneh & Balvardi, Mohammad & Akhavan, Hamid-Reza. (2021). Optimization of Ethanol-Assisted Aqueous Oil Extraction from a Cicadidae Sp. 10.1101/2021.01.24.427958. Link.
  3. Sanborn, Allen & Villet, Martin. (2021). A New Species of Buyisa Distant, 1907 (Hemiptera: Cicadidae: Cicadettinae: Cicadettini) from South Africa. Proceedings of the Entomological Society of Washington. 123. 10.4289/0013-8797.123.1.199. Link.
  4. Park, Gunhyuk & Moon, Byeong & Ryu, Seung & Kim, Wook & Lim, Hye-Sun. (2021). Cicadidae Periostracum Attenuates Atopic Dermatitis Symptoms and Pathology via the Regulation of NLRP3 Inflammasome Activation. Oxidative Medicine and Cellular Longevity. 2021. 1-16. 10.1155/2021/8878153. Link.
  5. Hill, Kathy & Marshall, David & Marathe, Kiran & Moulds, Max & Lee, Young June & Pham, Thai & Mohagan, Alma & Sarkar, Vivek & Price, Benjamin & Duffels, J. & Schouten, Marieke & Boer, Arnold & Kunte, Krushnamegh & Simon, Chris. (2021). The molecular systematics and diversification of a taxonomically unstable group of Asian cicada tribes related to Cicadini Latreille, 1802 (Hemiptera : Cicadidae). Invertebrate Systematics. 35. 10.1071/IS20079. Link.
  6. Allen F. Sanborn “A New Genus and Species of Neotropical Taphurini Distant, 1905 (Hemiptera: Cicadoidea: Cicadidae: Cicadettinae) from Brazil with a Note on the Taxonomic Position of Prosotettix Jacobi, 1907,” Proceedings of the Entomological Society of Washington, 123(1), 190-198, (29 January 2021). Link.
  7. Sarkar, Vivek & Mahapatra, Cuckoo & Mohapatra, Pratyush & Kunte, Krushnamegh. (2021). Description of three new species of the genus Mata Distant, 1906 (Hemiptera: Cicadidae: Cicadinae: Oncotympanini) with notes on their natural history from Indian state of Meghalaya, India. Zootaxa. 4908. 1-28. 10.11646/zootaxa.4908.1.1. Link.
  8. Kye, Geunho & Machta, Jonathan & Abbott, Karen & Hastings, Alan & Huffmyer, William & Ji, Fang & Liebhold, Andrew & Blackwood, Julie. (2021). Sharp boundary formation and invasion between spatially adjacent periodical cicada broods. Journal of Theoretical Biology. 515. 110600. 10.1016/j.jtbi.2021.110600. Link.

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