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April 30, 2016

64 Degrees Fahrenheit Eight Inches Deep

Filed under: James Edward Heath,Magicicada,Periodical — Dan @ 1:22 am

hot under the exoskeleton

Soil temperature triggers periodical cicada emergences:

James Edward Heath in his paper Thermal Synchronization of Emergence in Periodical “17-year” Cicadas (Homoptera, Cicadidae, Magicicada)1 discovered that periodical cicadas will emerge, on average, when the soil 8 inches below the ground reaches 64 degrees Fahrenheit:

Soil temperature at 20-cm [7.87 in] depth in seven locations averaged 17.89 C [64.202 F] at the time of emergence, regardless of date. Cicadas emerging from burrows had average body temperatures of 18.04 C [64.472 F]. Synchrony in emergence may be due to animals reaching a critical threshold temperature.

The soil warms their bodies and that triggers the cicadas to emerge.

A warm rain, which will seep into the earth and warm the bodies of cicadas, can trigger a “particularly intensive” emergence.

Alexander and Moore (1962) noted that emergences were par- ticularly intensive following warm rains ( > 2 0 C ? ). I believe this suggests that the soil temperature probably was near emergence tem- perature and the heat transported by the water percolating into the ground warmed the soil sufficiently to raise soil temperature above the emergence threshold.

Some unanswered questions I have are: 1) how long does the temperature have to be 64°F+ – just a moment, or a certain number of hours, and 2) because we know not all cicadas emerge instantaneously, what is the maximum temperature that for certain will clear them from the soil?

This is an example of a temperature probe use to study the emergence temperature of cicadas. The probe in this picture is held by cicada researcher Gene Kritsky:
Temperature probe

Note that males typically emerge before females, and that the larger ‘decim species emerge before the smaller cassini species.

For a more modern, crowd-sourced study of this see the radiolab Cicada Tracker project.

I think I know what you’re thinking: “I don’t own a temperature probe, how can I guess when the temperature is 64°F 8” below the soil? Direct sunlight, air temperature and rain warms the soil. Southern facing land will warm sooner than northern facing land. Land in direct sunlight will warm faster than land in shade. Rainfall on a 80°+ day will quickly do the trick, but two weeks of temps in the 70°’s should work as well. So keep an eye on the 10-day weather forecast and watch for those days in the 70s, and especially the 80s.

Minimum Flight Temperature:
Their body temperature needs to be a little warmer than that to fly. Their minimum flight temperature (MFT) is 18-21°C / 65-70°F. The temperature varies depending on the Brood and species. They’ll need a few more degrees before they’re fully functional, and start singing and mating.

So, until their bodies are about 72°F (“room temperature”) they won’t be flying, singing and mating.

Maximum voluntary tolerance temperature:
Maximum voluntary tolerance temperature (MVT) for periodical cicadas is 31-34°C / 88-93°F, again depending on Brood and species. Maximum voluntary tolerance is the point at which cicadas seek shade and when thermoregulation takes precedence over other behaviors.

See Thermal responses of periodical cicadas: within and between brood parity (Hemiptera: Cicadidae: Magicicada spp.) and Thermoregulation by Endogenous Heat Production in Two South American Grass Dwelling Cicadas (Homoptera: Cicadidae: Proarna) for more information.

1 Thermal Synchronization of Emergence in Periodical “17-year” Cicadas (Homoptera, Cicadidae, Magicicada) by James Edward Heath, American Midland Naturalist, Vol. 80, No. 2. (Oct., 1968), pp. 440-448.

April 27, 2016

Please use the correct imagery for 17 year cicadas

Filed under: Identify,Magicicada,Periodical,Video — Dan @ 1:01 am

If you’re writing an article about the coming emergence of the 17-year periodical cicadas, please use the correct genus & species of cicadas.

The genus of all 17 year cicadas is Magicicada, and they are never green. The three species of 17-year cicadas are M. septendecim, M. cassini, and M. septendecula. They’re all black with orange wings and legs and red eyes (some exceptions, but they’re never green).

An adult Magicicada septendecim by Dan Mozgai/cicadamania.com:

Brood II Magicicada septendecim

A newly emerged, teneral, Magicicada septendecim by Dan Mozgai/cicadamania.com:

Imagining Brood II Magicicada septendecim

A mating pair of Magicicada septendecim by Dan Mozgai/cicadamania.com:

A pair of mating Magicicada septendecims found in Woodbridge Township NJ

17-year cicada video:

A singing Magicicada septendecim:

Singing Magicicada septendecim from Cicada Mania on Vimeo.

A Magicicada septendecim laying eggs:

Magicicada septendecim ovipositing from Cicada Mania on Vimeo.

A Magicicada septendecim up close (deceased):

Close up of a tymbal of a Magicicada septendecim from Cicada Mania on Vimeo.

Magicicada on a tree (mostly Magicicada cassini):

For the sake of cicada correctness, feel free to use them in your article. Just credit cicadamania.com.

More photos: All the Magicicada galleries on the site.

If you are looking to license Magicicada images or HD Video, Roy Troutman has plenty of both. Reach out to him if interested. His images and video are tagged throughout the site.

Hundreds of shed cicada skins (exuvia) by Troutman:

2014 Ohio Exuvia Pile by Roy Troutman

Wrong Cicadas:

If the cicada you use in your article is green, it isn’t a 17-year cicada. I repeat: if the cicada is green it is not a 17-year cicada.

The cicada at the top of the Wikipedia page for cicadas is not a 17-year cicada, it’s an annual cicada called Neotibicen linnei:

Tibicen linnei
(photo credit for this Neotibicen linnei).

The cicada shedding its skin on a roll of paper towel… that’s not a 17-year cicada either:

Wikipedia
(Photo Credits for the molting cicada).

Looking for people to speak at a conference or “cicadacon”?

Need a speaker for a Cicada Convention or a Periodical Cicada Event? Try these folks:

April 15, 2016

Brood V 17-Year Cicadas Due in Spring of 2016

Filed under: Brood V,Magicicada,Periodical — Dan @ 1:50 am

Brood V 2016Brood V (5) 17-year cicadas will emerge in the spring of 2016 in Maryland, New York, Ohio, Pennsylvania, Virginia, and West Virginia.

Latest Updates:
(4/27/16): many sightings of nymphs are appearing on the Magicicada.org map. Someone shared a photo on Facebook of a nymph probably dug up by a rodent, like a vole.

(4/22/16): the Report a Cicada Sighting form is up on Magicicada.org.

FYI: periodical cicadas will not emerge in 2016 in Baltimore, Washington D.C. and New York City.

(4/2/16): Cleveland Metroparks posted a photo of a Magicicada nymph close to the surface.

About Brood V:

The cicada species that will emerge are Magicicada cassinii (Fisher, 1852), Magicicada septendecim (Linnaeus, 1758), and Magicicada septendecula Alexander and Moore, 1962. These periodical cicadas have a 17-year life cycle. The last time they emerged was 1999.

When: Generally speaking, these cicadas will begin to emerge when the soil 8″ beneath the ground reaches 64 degrees Fahrenheit. A nice, warm rain will often trigger a emergence. So, definitely May, but something might happen in April if we have a particularly hot spring.

Locations where they are likely to emerge:

This data comes from the Cicada Central Magicicada Database and other sources.

Although the cicadas will emerge in MD, NY, OH, PA, VA and WV, the area is limited and patchy. No Brood V cicadas for D.C., Cincinnati, or NYC (people have asked). Their range is closer to this map (with cicadas in the orange areas):

Brood V range

Maryland:

Counties: Garrett.

New York:

Specific locations in L.I.:

  • Wildwood State Park (maybe not… TDB)

Counties: Suffolk (Long Island).

Ohio:

Specific locations in Ohio:

  • The emergence should be good in the south eastern part of the state and in Summit, Medina, and southern Cuyahoga counties1.
  • Hocking State Forest, Hocking county, which is where James Edward Heath performed his investigation of periodical cicada Thermal Synchronization2.
  • Tar Hollow State Forest, in Laurelville, Hocking County, Ohio.
  • Strouds Run State Park, in Canaan Township, Athens County.
  • Athens, Athens County, Ohio
  • Findley State Park, Lorain County, Ohio.

Counties: Ashland, Ashtabula, Athens, Belmont, Carroll, Columbiana, Coshocton, Crawford, Cuyahoga, Fairfield, Franklin, Gallia, Guernsey, Harrison, Hocking, Jackson, Jefferson, Knox, Lake, Lawrence, Licking, Lorain, Mahoning, Medina, Meigs, Muskingum, Noble, Ottawa, Perry, Pickaway, Pike, Portage, Richland, Ross, Scioto, Seneca, Stark, Summit, Trumbull, Tuscarawas, Vinton, Washington, Wayne

Thanks to Roy Troutman, John Cooley, Chris Simon and Gene Kritsky for the tips!

Pennsylvania:

Counties:Allegheny, Fayette, Greene, Washington, Westmoreland

Virginia:

Specific locations in Virginia:

  • Douthat State Park, in Bath & Allegheny County Virginia.

Counties: Allegheny, Augusta, Bath, Highland, Richmond, Rockingham, Shenandoah

West Virginia:

Counties: Barbour, Boone, Braxton, Brooke, Cabell, Calhoun, Clay, Doddridge, Fayette, Gilmer, Grant, Greenbrier, Hampshire, Hancock, Hardy, Harrison, Jackson, Kanawha, Lewis, Marion, Marshall, Mason, Monongalia, Nicholas, Ohio, Pendleton, Pocahontas, Preston, Putnam, Raleigh, Randolph, Ritchie, Roane, Taylor, Tyler, Upshur, Webster, Wetzel, Wood

Learn more about Brood V:

Learn More:

Watch a cicada emerge from its skin

A whole lot of cicada nymphs

1 Kritsky, G., J. Smith, and N. T. Gallagher. 1999. The 1999 emergence of the periodical cicada in Ohio (Homoptera: Cicadidae: Magicicada spp. Brood V). Ohio Biological Survey Notes 2:43-47.

2 Thermal Synchronization of Emergence in Periodical “17-year” Cicadas (Homoptera, Cicadidae, Magicicada) by James Edward Heath, American Midland Naturalist, Vol. 80, No. 2. (Oct., 1968), pp. 440-448.

* The map is based on this map from the Wikimedia Commons by Lokal_Profil.

February 7, 2016

The Periodical Cicada Brood VII Revisited

Filed under: Brood VII,Magicicada,Periodical — Dan @ 9:22 pm

The Internet Archive has a lot of cicada documents and information, including a growing collection of articles from journals.

Today I came across a paper about Brood VII called The Periodical Cicada Brood vii Revisited (Homoptera, Cicadidae) by L. L. Pechuman, published in 1985 in the journal Entomology News (link to the article). Brood VII will be back in New York in 2018 (not too far away) so I’m glad I found this now. Brood VII is interesting because it is geographically isolated from other broods, near the Finger Lakes area of New York. This always makes me wonder what happened that led to their isolation (glacial melting, a massive die off of host trees… who knows).

The article is a quick, but melancholy read — unfortunately Brood VII is a small and dwindling brood; it has gone extinct in many areas, and has suffered over-predation by birds in recent emergences. “Populations were just not high enough to support ‘predator satiation'”, according to L. L. Pechuman.

People who witness massive periodical cicada emergences would never think that they were a fragile insect, but they are and papers like this make that fact abundantly clear.

November 24, 2015

Cicada Fun with Google Trends

Filed under: Australia,Brood X,Japan,Periodical,United States — Dan @ 7:59 pm

This article was inspired by Serious Fun with Google Trends by Simon Leather.

Google Trends is a Google website that lets you see trends in the search terms over time. When people search for “cicada” it usually means cicadas have emerged in their area at the time they search.

The following graph shows when people searched for “cicada” over the past 10 years in the United States. The largest spike, in May of 2004, coincided with the emergence of Brood X.

2004-2015

You might think that periodical cicada emergences cause the largest spikes, but not always — and not just because periodical cicadas don’t emerge every year.

2004: Cicada searches spiked May 16-22, which was Brood X – Magicicadas.
2005: Jul 31-Aug 6 spike which was for Neotibicen Cicadas. No periodical cicadas.
2006: Aug 13-19, Neotibicen Cicadas. No periodical cicadas.
2007: May 20-26, Brood XIII – Magicicadas.
2008: Brood XIV Magicicadas emerged (spike Jun 8-14), but the largest spike was Jul 29-Aug 2, Neotibicen Cicadas.
2009: Aug 16-22, Neotibicen Cicadas.
2010: Aug 8-14, Neotibicen Cicadas.
2011: May 29-Jun 4, Brood XIX – Magicicadas.
2012: Jul 29-Aug 4, Neotibicen Cicadas.
2013: May 5-11, Brood II – Magicicadas.
2014: Brood XXII – Magicicadas had a relatively small spike May 25-31, compared with Aug 24-30 for Neotibicen Cicadas (late season due to cool weather). There was also a teeny bit of a spike around January of 2014 due to the “cicada 3301” meme/game.
2015: Brood XXIII & IV Magicicadas emerged (spike around Jun 7-13), but the largest spike was around Aug 9-15 for Neotibicen Cicadas.

Which cities had the most cicada searches over the past 10 years? Cincinnati, Omaha, Nashville, Baltimore, Washington, Chicago, Alexandria, St. Louis, Philadelphia, and Charlotte. Time for me to move to Cincinnati!

How about Australia? Cicada searches in Australia spike in December. The largest spike was in December of 2013, which was indeed a big year for cicada emergences in Australia.

The variation year after year suggests that there is a degree of periodicity to Australian cicadas. There is always a spike around December, but some years see bigger spikes than others. Just speculation (partially because the 2004 & 2005 data seems absent) but perhaps there is a 9 or 11 year proto-periocity happening.

In terms of cities, Sydney has had the most cicada searches:

How about Japan?! August is the best month for cicadas (セミ) in Japan.

Yokohama, Chiba & Saitama generates the most cicada searches:

October 11, 2015

Look/Listen for Brood IX Cicada Stragglers in 2016

Filed under: Brood IX,Magicicada,Periodical Stragglers — Dan @ 11:11 am

There is a high probability that Brood IX (17 Year Magicicada) stragglers will emerge in 2016. Look for them in southern West Virginia, western Virginia and north-west North Carolina:

Brood IX

M. septendecim, M. cassini, and M. septendecula are all part of this brood.

Learn more about periodical cicada stragglers.

Visit Magicicada.org’s Brood IX page for detailed information.

July 14, 2015

Annual Cicada Mania

Filed under: Annual — Dan @ 1:01 am

The 2015 annual cicada season is well underway.

The annual cicada species are out now (now means May to September) around the United States. Not all annual cicadas are out yet — some won’t arrive until late summer — but many are currently out in the southernmost states, and will soon arrive in northern states.

Wonder which annual cicadas are in your area? Try Biogeography of the Cicadas (Hemiptera: Cicadidae) of North America, North of Mexico, a PDF that features maps of annual cicadas. Also try our new U.S.A. & Canada Cicada Search search tool.

Wonder what they look like? Start at our Cicadas by Genus and Species page.

Wonder what they sound like? Try Insect Singers.

Here are a small portion of the species that can be found in the USA:

Diceroprocta apache
Diceroprocta apache
Found in: AZ, CA, CO, NV, UT
Diceroprocta olympusa
Diceroprocta olympusa
Found in: AL, FL, GA, MS, NC, SC
Neocicada hieroglyphica
Neocicada hieroglyphica
Found in: AL, AR, DE, FL, GA, IL, IN, KS, KY, LA, MD, MS, MO, NJ, NY, NC, OH, OK, SC, TN, TX, VA
Okanagana bella
Okanagana bella
Found in: AB, AZ, BC, CA, CO, ID, MT, NV, NM, OR, SD, UT, WA, WY
Okanagana rimosa
Okanagana rimosa
Found in: AB, BC, CA, CT, ID, IL, IN, IA, ME, MB, MD, MA, MI, MN, MT, NV, NB, NH, NJ, NY, ND, OH, ON, OR, PA, QC, SD, UT, VT, VA, WA, WI, WY
Tibicen superbus
Tibicen superbus
Found in: AR, KS, LA, MO, NM, OK, TX
Tibicen dorsatus
Tibicen dorsatus
Found in: AR, CO, ID, IL, IA, KS, MO, MT, NE, NM, OK, SD, TX, WY
Cicadetta calliope
Cicadetta calliope
Found in: AL, AR, CO, FL, GA, IL, IN, IA, KS, KY, LA, MD, MS, MO, NE, NC, OH, OK, SC, SD, TN, TX, VA
Tibicen pruinosus
Tibicen pruinosus
Found in: AL, AR, CO, FL, GA, IL, IN, IA, KS, KY, LA, MA, MD, MI, MN, MS, MO, NE, NJ, NY, NC, OH, OK, PA, SC, SD, TN, TX, VA, WV, WI

July 12, 2015

Why do Magicicada stay underground for 13 or 17 years?

Filed under: FAQs,Life Cycle,Magicicada,Periodical — Dan @ 8:01 pm

People ask: why do periodical cicadas stay underground for 17 or 13 years?

There are three parts to this puzzle that people are interested in:

  1. How cicadas count the years as they go by.
  2. Why prime numbers? 13 and 17 are prime.
  3. Why is their life cycle so long? They are one of the longest living insects.

Cicadas likely don’t count like people do (“1,2,3,4…”) and you won’t find scratch marks inside the cell (where they live underground) of a Magicicada, marking off the years as they go by. However, there is a kind of counting going on, and a good paper to read on that topic is How 17-year cicadas keep track of time by Richard Karban, Carrie A. Black and Steven A. Weinbaum. (Ecology Letters, (2000) Q : 253-256). By altering the seasonal cycles of trees they were able to make Magicicada emerge early, proving that cicadas “count” seasonal cycles, perhaps by monitoring the flow and quality of xylem sap, and not the passage of real time.

Why prime numbers, and why is the life cycle so long? This topic fascinates people. The general consensus is that the long, prime numbered life-cycle makes it difficult for an above-ground animal predator to evolve to specifically predate them. Read Emergence of Prime Numbers as the Result of Evolutionary Strategy by Paulo R. A. Campos, Viviane M. de Oliveira, Ronaldo Giro, and Douglas S. Galva ̃o (PhysRevLett.93.098107) for more on this topic. An argument against that theory is that a fungus, Massospora cicadina, has evolved to attack periodical cicadas regardless of their life cycle. Of course, a fungus is not an animal. Maths are easy for fungi.

There are also questions about why there are 13 and 17 year life cycles, why a 4 year acceleration of a brood might occur1 and why Magicicada straggle.

1 This is a good place to start: Genetic Evidence For Assortative Mating Between 13-Year Cicadas And Sympatric”17-Year Cicadas With 13-Year Life Cycles” Provides Support For Allochronic Speciation by Chris Simon, et al, Evolution, 54(4), 2000, pp. 1326–1336.

30 Things Cicadas Do

Filed under: FAQs,Life Cycle — Dan @ 11:59 am

One of the most frequently asked questions we receive is: “what do cicadas do“? This question is similar to the question “what is the purpose of cicadas” — the answers to both questions help people understand why these fascinating, unusual creatures exist at all.

The simplest reduction of their life cycle is:
1) They hatch from an egg.
2) They burrow underground where they will drink from plant roots for most of their lives.
3) They leave the underground, and become adults.
4) The males make sounds that attract females.
5) Males & females court & mate.
6) Females lay fertilized eggs in the branches of plants, and the cycle continues.
7) They die.

The specifics of a cicada’s life cycle varies from species to species, but here is a more detailed view of what cicadas do:

From egg to 1st instar nymph:
1) Cicada nymphs hatch from eggs.

Cicada egg nests

2) Nymphs feed on plant fluids which they accesses thanks to the egg-nest groove made by their mothers.

3) They leave the groove, and drift to the ground. Their decent to the ground doesn’t hurt them because they weigh so little.

4) Once on the ground, they dig into the soil until they find small rootlets, from which they will feed.

Once Underground:
5) Underground, they will tunnel/dig
6) and establish a cell
7) from where they’ll comfortably feed. Cicadas feed on the xylem sap of plants. With the help of bacteria they transform the water, minerals and amino acids found in tree fluids to the tissues of their own bodies.
8) They pee, in fact they seem to use excess plant fluid to moisten soil to help mold the walls of their cells.
9) Throughout their life underground they will move from root to root… as plant root systems change with the seasons, when roots die off, or perhaps to avoid predators.
10) Underground, a cicada may (depending on the species) go through four instars, molting three times (see an image of the four instars).

Preparing to emerge:
11) Cicadas will build a tunnel to the surface of the ground, in preparation for their emergence.

12) Cicadas often take that a step further an build a chimney/turret above ground. This often happens in shady areas or when the ground is muddy.

Once above ground:

13) They emerge from their tunnels
14) Cicadas run as fast as they can…
15) And find a surface perpendicular to the ground, hold tight, and begin to molt…
16) During the molting process (ecdysis), cicadas perform many acrobatic moves to separate themselves from their nymphal skin, including pulling their old trachea from their bodies.
A Brood XIV Magicicada straggler, emerged 4 years late.
17) Once outside their nymphal skin, they will inflate their wings
18) … and expand various parts of their bodies, like their heads.
19) They will change color.
20) Once their bodies are hard enough (sclerotization counts as a thing they do)…
21) They will either seek shelter, perhaps by crawling up higher along a tree trunk…
22) or if your are a Magicicada, you might stick around in the hopes that a predator will eat you.

Mating and Reproduction:

23a) If you are a Male cicada, you are going to sing… unless you belong to a species that cannot sing, in which case, you’ll move your wings in a way that will produce a sounds.
There are many types of songs: a) distress calls, b) calls to establish territory, c) calls to attract females, d) including choruses of many cicadas, and e) courting calls
23b) Female cicadas, and some male cicadas, move their wings to make sounds, also in an effort to attract and engage a mate.
24) Most cicadas (aside from Magicicada during the early days of their adult lives) will try avoid being eaten by predators.
25) They’ll fly, of course.
26) Cicadas, like Magicicada, will establish chorusing centers, which are places where the male cicadas sing together and females come to meet them.
27) Male and female cicadas will court
28) and mate
29) the female cicada will lay her eggs in grooves (ovipositing) she etches into a suitable plant stem, and we’re back to step 1.

30) The last thing cicadas do, of course, is die, and return the nutrients found in their bodies to the soil, where they will be broken down and absorbed by the plants they fed upon.

Some things cicada do not do:

Here are some things cicadas do not do:

1) They don’t seek shelter during the fall months (i.e. they don’t try to live inside your house), unlike Ladybugs or Stinkbugs.
2) They don’t sting or otherwise pass venom onto people.
3) They don’t chew plant leaves, like caterpillars or grasshoppers.
4) They don’t dump garbage in the ocean.

July 5, 2015

What are Broods?

Filed under: FAQs,Magicicada,Periodical — Dan @ 6:29 pm

It is important to note that when we talk about cicada broods, we are talking about the 17 & 13 year periodical Magicicada cicadas. We are not talking about Tibicen or other species.

All Broods

There are 12 groups of Magicicadas with 17 year life cycles, and 3 groups of Magicicadas with 13 year life cycles. Each of these groups emerge in a specific series of years, rarely overlapping (17 & 13 year groups co-emerge every 221 years, for example). Each of these groups emerge in the same geographic area their parents emerged. These groups, each assigned a specific Roman numeral, are called broods.

Gene Kritsky’s book, Periodical Cicadas: The Plague and the Puzzle, documents the history of the recognition and naming of the broods. The first person to document that different groups of periodical cicadas emerged in different years was Nathaniel Potter in 1839. Benjamin D. Walsh and Charles V. Riley devised the system for numbering the different broods in 1868, and then C. L. Marlatt sorted the 17 year broods out from the 13 year broods, giving us the system we have today.

Visit our Broods page which features a grid of the Brood names, their life span, when & where they’ll emerge next and links to maps.


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